2009 in paleontology


Arthropods

Cephalopods

Three new species of extinct Octopoda discovered in 2009. The species – Keuppia hyperbolaris, Keuppia levante, and Styletoctopus annae – lived about 95 million years ago, and bear a strong resemblance to modern octopuses, suggesting that the Octopoda order has remained relatively unchanged for tens of millions of years. The fossils included evidence of arms, muscles, rows of suckers, ink, and internal gills. The discovery was made by a team led by Dirk Fuchs of the Freie University, which is located at Berlin, Germany. The fossils were found at Hakel and Hadjoula, Lebanon. Various new ammonoid taxa were named, including Ivoites.
NameNoveltyStatusAuthorsAgeUnitLocationNotesImages

Keuppia

Gen et sp nov

Valid

Fuchs, Bracchi, & Weis

Upper Cenomanian



Styletoctopus

gen et sp nov

Valid

Fuchs, Bracchi, & Weis

Upper Cenomanian


Ivoites''

gen nov

Valid

De Baets, Klug, & Korn

Lower Emsian

Cartilaginous fish

Bony fish

Newly named amphibians

NameStatusAuthorsAgeUnitLocationNotesImages

Baphetes orientalis

Valid

  • Milner
  • Milner
  • Walsh
Late Carboniferous
-

Cratia

Valid

  • Báez
  • Moura
  • Gómez
Lower Cretaceous
Crato Formation


Possible stem neobatrachian

Eurycephalella

Valid

  • Báez
  • Moura
  • Gómez
Lower Cretaceous
Crato Formation


A hyloid

Nannaroter

Valid

Early Permian

The smallest known ostodolepid microsaur

Nesovtriton

Valid

  • Skutschas
Turonian
Bissekty Formation


A cryptobranchoid salamander

Regalerpeton

Valid

Early Cretaceous
Huajiying Formation


A cryptobranchoid salamander

Spinarerpeton

Valid

  • Klembara
Early Permian
Boskovice Furrow


A discosauriscid seymouriamorph
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------
-------

Basal reptiles

Newly named basal reptiles

NameStatusAuthorsDiscovery yearAgeUnitLocationNotesImages

Australothyris

Valid

  • Modesto
  • Scott
  • Reisz
Middle Permian
Tapinocephalus Assemblage Zone


A basal parareptile
-

Procolina

Valid

  • Borsuk−Białynicka
  • Lubka
early Late Olenekian
Czatkowice 1


A procolophonine procolophonid
-

Turtles

Newly named turtles

NameStatusAuthorsDiscovery yearAgeUnitLocationNotesImages

Angolachelys

Valid

Turonian

Aurorachelys

Valid

  • Vandermark
  • Tarduno
  • et al.
Late Cretaceous
Strand Fiord Formation


Basilochelys

Valid

  • Tong
  • Claude
  • et al.
Late Jurassic/Lower Cretaceous
Phu Kradung Formation


Cedrobaena

Valid

  • Lyson
  • Joyce
Paleocene
Fort Union Formation

USA

New genus for "Plesiobaena" putorius Gaffney, 1972

Chelonoidis alburyorum

Valid

  • Franz
  • Franz
Holocene

A tortoise.

Chinlechelys

Valid

  • Joyce et al.
Late Triassic
Bull Canyon Formation


A basal member of Testudinata. The type species is C. tenertesta. The genus Chinlechelys was considered to be a junior synonym of the genus Proganochelys by Joyce, though the author maintained C. tenertesta as a distinct species within the latter genus.

Derrisemys

Junior synonym

  • Hutchison
Early Paleocene
USA

Junior synonym of Hutchemys.

Eileanchelys

Valid

  • Anquetin
  • Barrett
  • et al.
Middle Jurassic
Kilmaluag Formation


Hutchemys

Valid

  • Joyce
  • Revan
  • Lyson
  • Danilov
Paleocene
Fort Union Formation
Tullock Formation

USA

A plastomenine softshell turtles

Kinkonychelys

Valid

Late Cretaceous
Maevarano Formation


Palatobaena cohen

Valid

  • Lyson
  • Joyce
Maastrichtian
Hell Creek Formation

USA

A baenid

Peckemys

Valid

  • Lyson
  • Joyce
Late Cretaceous
Hell Creek Formation

USA

A baenid

Plastomenoides

Junior synonym

  • Hutchison
Early Paleocene
USA

Junior synonym of Hutchemys.
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------
--------

Archosauromorphs

Basal archosauromorphs

Archosaurs

Lepidosauromorphs

Basal lepidosauromorphs

Plesiosaurs

Synapsids

Non-mammalian

NameStatusAuthorsAgeUnitLocationNotesImages

Protuberum

Valid

  • Reichel
  • Schultz
  • Soares
Middle Triassic
Santa Maria Formation


Raranimus

Valid

  • Liu
  • Rubidge
  • Li
Middle Permian
Xidagou Formation



Yuanotherium

Valid

  • Hu
  • Meng
  • Clark
Late Jurassic
Shishugou Formation

Mammals

Plants

Angiosperms

Relevant research in other sciences

Evolutionary biology

A study noting the effects of the KT mass extinction on Earth's modern biota is published.

Geology

Complete author list

As science becomes more collaborative, papers with large numbers of authors are becoming more common. To prevent the deformation of the tables, these footnotes list the contributors to papers that erect new genera and have many authors.