The Wing's first predecessor is the 5th Combat Crew Replacement Center Group, which was organized under VIII Composite Command at High Wycombe, England, the headquarters of VII Bomber Command on 5 November 1943. The following February, it moved to RAF Cheddington, England, with the mission of training newly arriving replacement bomber crews. The group moved to RAF Greencastle, Northern Ireland, in July 1944, and was disbanded there on 5 August 1944.
Missile training
activated the 705th Strategic Missile Wing on 20 February 1958 at RAF Lakenheath, England, to monitor the PGM-17 Thor IRBM program in the United Kingdom and provide technical assistance to the four Royal Air Force Thor squadrons. The wing never became operational, and its only manning was by additional duty personnel of the 7th Air Division Directorate of Ballistic Missiles and six weeks after activation, the wing moved to South Ruislip Air Station, the division's station, where it remained through 1960. Even its commanding officer, Col. William R. Yancy acted in that capacity as an additional duty from March to November 1958, The wing's 672d Technical Training Squadron was located at RAF Feltwell, where, on 22 June 1959, No. 77 Squadron RAF became the first British-based Thor squadron to reach operational status. The 672d moved to RAF Lakenheath, from which location it trained the remaining three RAF missile squadrons. On 11 Septemberand 22 December 1959, the second and third British-based Thor squadrons were declared operational and assigned to Royal Air Force personnel. When SAC turned over the fourth and final British-based Thor to the Royal Air Force on 22 April 1960, the deployment of the Thor IRBM weapon system in the United Kingdom was completed. The wing was inactivated on 1 April 1960, followed shortly by the elimination of 7th Air Division's Directorate of Ballistic Missiles. While the missiles were assigned to the RAF for control, SAC retained control over the squadron's nuclear warheads and assigned detachments of the attached 99th Aviation Depot Squadron with each RAF missile squadron to maintain custody and control over, and maintain reentry vehicles and warheads until it received U. S. warhead release orders. It also operated USAF communications facilities and trained Royal Air Force personnel on Thor missile operations.
Lineage
; 5th Combat Crew Replacement Center Group
Constituted as the 5th Combat Crew Replacement Center Group
Reconstituted on 31 July 1985 and consolidated with the 705th Strategic Missile Wing as the 510th Tactical Missile Wing
; 705th Strategic Missile Wing
Constituted as the 705th Strategic Missile Wing on 6 February 1958
Consolidated with the 5th Combat Crew Replacement Center Group on 31 July 1985 as the 510th Tactical Missile Wing
; 510th Tactical Missile Wing
5th Combat Crew Replacement Center Group and 705th Strategic Missile Wing consolidated on 31 July 1985 as the 510th Tactical Missile Wing