Aquilinae
The Aquilinae are a subfamily of eagles of the family Accipitridae. The general common name used for members of this subfamily is "booted eagle", although this is also the common name of a member of the subfamily. At one point, this subfamily was considered inclusive with the Buteoninae based probably on some shared morphological characteristics. However, research on the DNA of the booted eagles has shown that they are a monophyletic group that probably have had millions of years of separation from other extant forms of accipitrid.
Description
The Aquilinae consists of medium-sized to very large species. Modern species range in mass from the recently recognized pygmy eagle of Papua New Guinea, which weighs about to the martial eagle and golden eagle, which both weigh about on average. In wingspan, extant Aquilinae range from the pygmy eagle, with a median of, to the wedge-tailed eagle and martial eagles, which average about and in wingspan, respectively. Total length can vary from, in the pygmy and wedge-tailed eagles, respectively. The record sizes for wild booted eagles are for a golden eagle in body mass and for a wedge-tailed eagle in wingspan. Dwarfing these species, the booted eagle species Haast's eagle of New Zealand, was probably the most massive species ever known in the entire accipitrid family, with females averaging an estimated and perhaps weighing up to, but went extinct by the 16th century when human colonizers killed off their prey. Beyond their typically large size, Aquilinae species have few outward shared characteristics as they are a fairly diverse subfamily. Nonetheless, every species shares the feature of their legs being covered in feathers. Only two buteonine hawks share this feature beyond the Aquilinae, in a presumed case of convergent evolution. Many species within the subfamily are colored for camouflage with varying patterns of brown, black, yellowish or white being commonplace, darker colors being favored by forest dwelling varieties and brown to straw coloring common to open country species. A few more brightly colored species, with striking patterns of chestnut in about four, none more striking than the ornate hawk-eagle. Roughly half of booted eagle species have a strikingly different looking juvenile plumage, which can in some species be nearly all white, but even where the differences are subtle between adults and immatures, usually some physical differences are noticeable such as white wing patches in traditional Aquila species, to visually distinguish young from adults. Many species have a notable head-crest, such as Nisaetus and most Spizaetus, with the feature being most extreme in the long-crested eagle. Like most accipitrids, they have large powerful hooked beaks for tearing flesh from their prey, strong legs and powerful talons. Compared to other large raptors, the feet and talons of booted eagle species are often particularly large and powerful relative to their size. The most extreme example seems to be the crowned eagle, which has a hallux-claw of around about the same sized hallux-claw as the Philippine eagle , which weighs on average nearly twice as much. The booted eagles also have extremely keen eyesight to enable them to spot potential prey from a distance.Life histories and range
Booted eagles are varying in their habitats and habits, being found on every continent inhabited by accipitrids, which includes Antarctica. They may inhabit all of the world's terrestrial habitats, with the majority of species being largely forest dwelling, but several preferring all kinds of open habitat from steppe to prairie to tundra as well as most rocky or mountainous areas. By far the widest ranging species of booted eagle is the golden eagle, which is distributed in all of North America and much of Eurasia including a majority of Europe, often along most mountainous terrains therein but also other land-based habitats with semi-open or hilly hunting grounds accessible. So-called "hawk-eagles" are forest dwelling booted eagle types largely represented by Spizaetus and Nisaetus. Despite many similar features in the two main hawk-eagle genera, they apparently evolved separately for some time and are not closely related. The peak diversity for Aquilinae are in Africa, where nearly half of extant species are known to dwell at least seasonally, exploiting a diversity of different hunting styles, habitats, nesting habits and general body forms. All booted eagle species are opportunistic predators which prey mostly on small vertebrates, often subsisting on a wide range of mammals, birds, reptiles and amphibians as they become locally available or abundant. Species favoring more open habitats tend to hunt most often on the wing, using distance or hugging contours of the ground to surprise prey. Forest dwelling species are more likely to be perch-hunters, watching for prey activity from a high tree perch, but every kind of booted eagle can vary their hunting techniques when necessary. In cases where they become specialists, booted eagles tend to be mammal hunters, such as the considerable dependence of Spanish imperial eagle on rabbits, of breeding steppe eagle on sousliks or Verreaux's eagle on hyraxes. Exceptions include the Ayres's hawk-eagle and, to a lesser extent, the rufous-bellied eagle, both of which are smallish, swift-flying bird-hunting specialists who dive on woodland birds while the black eagle feeds largely on the contents of other birds' nests. Most booted eagle species have relatively large feet and talons and can semi-regularly go after prey as large or larger than themselves, even smaller species can occasionally dispatch prey of up to two to three times their own weight. Rare attacks on large prey, such as ungulates, adult monkeys or medium-sized carnivores or other much larger animals, usually involve the larger species of booted eagle, whereas such attacks are exceptionally rare to non-existent in eagles of other lineages. At least four large booted eagles have been known to dispatch prey weighing or more..
Booted eagles, like almost all raptorial birds, are strongly territorial and tend to maintain expansive home ranges against conspecifics. Given their need for ample hunting ranges, the territory of most booted eagles tend to be extremely large, with several dozen square kilometers being common. Territories are usually maintained with display flights but vocal displays are known in some of the denser forest dwelling varieties. Nests of Aquilinae tend to be typically large as in most eagles, with ample surface space needed, whether located in trees, rocky formations or on the ground. As is typical in many raptorial birds, pair bonds are strong between males and females and in many booted eagle species, they may mate for life. Primarily females incubate the egg and brood the young, while males usually have the responsibility of food capture. Clutch sizes are usually small in booted eagles, rarely more than 3 eggs are laid, and most parents will manage to produce only one to two fledglings depending on species. In most booted eagles, beyond threats from the outside world, most eggs are laid and hatched at intervals, thus one sibling is usually considerably bigger and often kills its younger siblings. While brood size and fledgling rates are typically low, booted eagles may have even lower productivity due to the long dependence period of the young relying on their parents for food and protection, and some species may only nest every other year.
General status
Due to their large territories and low productivity, most booted eagle species are sparsely distributed and not infrequently uncommon-to-rare even in regional strongholds where ample habitat remains. Booted eagle species tend to be highly sensitive to human activities, mainly habitat alteration or destruction, human disturbance, collision with man made objects, accidental or intentional killing of staple prey species and various forms of persecution by humans. Due mainly to these factors, seven species of booted eagle are currently classified as Vulnerable to extinction by the IUCN. Four little-known forest dwelling species are classified in more severe status as Endangered: the Philippine hawk-eagle, Pinsker's hawk-eagle, Javan hawk-eagle and the black-and-chestnut eagle. A very different eagle, the steppe eagle, is also considered Endangered despite having been considered one of the most numerous of all eagles after a disastrous, ongoing decline mostly due to electrocutions from dangerous powerlines, poisonings and increasing steppe fires around nests, these killing them off en mass while breeding and migrating. An even more drastic classification of Critically Endangered was given to the Flores hawk-eagle. In all five endangered hawk-eagles, near epidemic levels of forest habitat degradation has been determined as the primary cause of their declines.Systematics
SUBFAMILY AQUILINAE| Image | Genus | Living Species |
| Stephanoaetus Sclater, 1922 |
| |
| Nisaetus Hodgson, 1836 | ||
| Lophotriorchis Sharpe, 1874 | ||
| Polemaetus Heine, 1890 | ||
| Spizaetus Vieillot, 1816 | ||
| Ictinaetus Blyth, 1843 | ||
| Lophaetus Kaup, 1847 | ||
| Clanga Adamowicz, 1858 | ||
| Aquila Brisson, 1760 | ||
| Hieraaetus Kaup, 1844 |