Aspleniineae


Aspleniineae is a suborder of ferns in the order Polypodiales. It is equivalent to the clade eupolypods II in earlier systems; it is also treated as a single very broadly defined family Aspleniaceae. The suborder generally corresponds with the order Blechnales as described by J. L. Reveal in 1993. Aspleniineae includes some important ferns, including Onoclea sensibilis, the sensitive fern, which grows as a virtual weed throughout much of its temperate North American range, and ferns of the genus Thelypteris, a genus that has shown remarkable speciation. It also includes one of the more common horticultural ferns, Matteuccia struthiopteris, the ostrich fern.

Taxonomy

In the Pteridophyte Phylogeny Group classification of 2016, the group is treated as the suborder Aspleniinae, and divided into 11 families. Alternatively, it may be treated as a single, very broadly circumscribed family Aspleniaceae sensu lato, which is then divided into subfamilies. The relationship between the two approaches is shown in the table below.
PPG IChristenhusz & Chase
Suborder Aspleniineae H.Schneid. & C.J.RothfFamily Aspleniaceae Newman
Family Cystopteridaceae Shmakov Subfamily Cystopteridoideae Ching & Z.R.Wang
Family Rhachidosoraceae X.C.Zhang Subfamily Rhachidosoroideae M.L.Wang & Y.T.Hsieh
Family Diplaziopsidaceae X.C.Zhang & Christenh. Subfamily Diplaziopsidoideae Christenh.
Family Desmophlebiaceae Mynssen
Family Hemidictyaceae Christenh. & H.Schneid. Subfamily Asplenioideae Link
Family Aspleniaceae Newman Subfamily Asplenioideae Link
Family Woodsiaceae Herter Subfamily Woodsioideae Schmakov
Family Onocleaceae Pic.Serm. Subfamily Blechnoideae Hook.
Family Blechnaceae Newman Subfamily Blechnoideae Hook.
Family Athyriaceae Alston Subfamily Athyrioideae B.K.Nayar
Family Thelypteridaceae Ching ex Pic.Serm. Subfamily Thelypteridoideae C.F.Reed

Phylogenic relationships

The following diagram shows a likely phylogenic relationship between the families of Aspleniineae, based on Lehtonen , and Rothfels & al..