Boris Schwanwitsch Boris Nikolayevich Schwanwitsch , Борис Николаевич Шванвич, was a Russian entomologist who specialised in Lepidoptera . He is best known for his studies of the colour pattern of the wings . Boris Schwanwitz graduated from the St. Petersburg University. After graduation, he changed a number of academic positions: assistant lecturer in Entomology at the Stebut Agricultural School , assistant lecturer and private-docent at Petrograd University , professor at the Perm University . In 1930, he returned to Leningrad to take the position of the head of Entomology department of the Leningrad University Vice-president of the Entomological Society of USSR and the chair of the Zoology section of the Leningrad Naturalists Society. In a series of papers he reconstructed the groundplan of the colour-pattern of the wings, first for the Rhopalocera , then for Heteroceran families. He formulated the stereomorphism principle, according to which the cryptic effect of the colour pattern is a result of its 'flattening' or 'disjunctive' effect. To prove his point he built the plaster three-dimensional models of the lepidopteran wings, the photographs of which looked like an actual colour pattern of stripes and shades. Among his other important contributions are a textbook in entomology with a large morphology section heavily based on Snodgrass and Weber , and a book on practical apiculture. His gravestone is ornamented with a reproduction of the groundplan of the colour-pattern of lepidopteran wings from his textbook.Selected publications On the groundplan of the wing-pattern in nymphalids and certain other families of rhopalocerous Lepidoptera. Proceedings of the Zoological Society of London , ser. B, 34: 509-528. Wing pattern in Papilionid Lepidoptera Entomologist vol. 76
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