Coryphodon


Coryphodon is an extinct genus of mammal.
Coryphodon was a pantodont, a member of the world's first group of large browsing mammals. It migrated across what is now northern North America, replacing Barylambda, an earlier pantodont. It is regarded as the ancestor of the genus Hypercoryphodon of Late Eocene Mongolia.
Coryphodon is known from many specimens in North America and considerably fewer in Europe, Mongolia, and China. It is a small to medium-sized coryphodontid who differs from other members of the family in dental characteristics.

Description

At about at shoulder height and in body length, Coryphodon is one of the largest-known mammals of its time. The creature was very slow, with long upper limbs and short lower limbs, which were needed to support its weight. Coryphodon does not seem to have been in need of much in the way of defences, however, since most known predators of the time seem to have been much smaller than Coryphodon.
Coryphodon had one of the smallest brain/body ratios of any mammal, living or extinct, possessing a brain weighing just and a body weight of around.
Estimates of Coryphodon's body mass have varied considerably. Based on a regression analysis of ungulates, estimated the mean body mass for the type species C. eocaenus to, for C. radians, and possibly as much as for C. proterus and C. lobatus.

Taxonomy and systematics

Since the first fossil was found in Wyoming, the taxonomy of Coryphodon and its family have been in disarray — five described genera have been synonymized with Coryphodon and thirty-five proposed species have been declared invalid.

Species

Coryphodon evolved from the Late Paleocene C. proterus, one of the largest species found and the only one known from the Clarkforkian NALMA. The body size then decreased until C. eocaenus appears at the Clarkforkian-Wasatchian transition, from where Coryphodon evolved into the large species C. radians. C. radians in its turn evolved into two contemporaneous species that appear in the Early Eocene, the small C. armatus and the very large C. lobatus. These changes in size are thought to be linked to global climate change, with the size minimum in the Coryphodon lineage occurring shortly after Paleocene-Eocene boundary.

Paleobiology

Feeding and diet

Coryphodon had a semi-aquatic lifestyle, likely living in swamps and marshes like a hippopotamus, although it was not closely related to modern hippos or any other animal known today. Coryphodon had very strong neck muscles and short tusks that were probably used to uproot swamp plants. The other teeth in the mouth were suited for processing plants that had been grabbed by browsing.
Fossils found on Ellesmere Island, near Greenland, show that Coryphodon once lived there in warm swamp forests of huge trees, similar to the modern cypress swamps of the American South. Though the climate of the Eocene was much warmer than today, plants and animals living north of the Arctic Circle still experienced months of complete darkness and 24-hour summer days. Isotopic studies of tooth enamel revealed that during the summer period of extended daylight Coryphodon would eat soft vegetation such as flowering plants, aquatic plants and leaves. However during the extended periods of darkness when plant photosynthesis was impossible, Coryphodon would switch to a diet of leaf litter, twigs, evergreen needles and most revealingly fungi, an organism and food source that does not require light to grow. Not only does this study reveal the dietary range of Coryphodon, but it also reveals the behaviour of the northern populations living within the Arctic Circle. In this respect, Coryphodon did not migrate south or hibernate, it simply switched between two seasonal food sources.

Sexual dimorphism

noticed a sexual dimorphism in Coryphodon: the canines tend to be either very large or very small compared to cheek teeth, and, comparing to modern hippos, there is reason to assume males had larger canines than females.