Doswell Formation
The Doswell Formation is a geologic unit of Upper Triassic age, part of the Newark Supergroup. The Doswell Formation was originally named to refer to a geological sequence which forms the lower part of the sedimentary fill of the Taylorsville Basin in Virginia and Maryland. This sequence was deposited by lakes and rivers in the developing rift basin. However, a 2016 study determined that several geological layers in Pennsylvania as well as the neighboring Richmond Basin of Virginia also qualified as components of the Doswell Formation.
The most diverse and fossiliferous component of the Doswell formation is the Vinita member, also sometimes referred to as the Turkey Branch, Tuckahoe, or Falling Creek Formations in earlier publications. The Doswell formation is biostratigraphically characterized by a fauna including the fish Dictyopyge macrurus and the conchostracan Laxitextella multireticulata. The Richmond Basin has several notable fossil sites, such as the Tomahawk site which has hundreds of fossils from the cynodont Boreogomphodon, and the Winterpock site which has an extraordinarily diverse assortment of plants. The Taylorsville Basin is much more restricted in terms of fossil locales, but it was home to the unusual armored reptile Doswellia.
The Doswell Formation is among the oldest Triassic formations on the east coast of the United States. It is believed to belong to the early Carnian age of the Triassic based on its fauna and flora, which is distinctly dominated by tetrapods similar to gondwanan groups, as well as a high diversity of humidity-loving plants such as ferns and cycads. This gives it a distinct disconnect from the younger formations of the Newark Supergroup, which typically have a fauna similar to the Triassic formations of the western United States and a flora including elements such as conifer trees, which are better adapted for drier conditions.
History and stratigraphy
Weems (1980)'s interpretation
The Doswell Formation was originally described by USGS paleontologist Robert E. Weems in 1980. He used it to refer to the entire Triassic geological sequence preserved at the Taylorsville basin, overlying the much older Carboniferous "Petersburg granite" which predated the basin, and underlying much younger Cretaceous and Cenozoic gravel. Weems subdivided the formation into several "members". The oldest of these was the Stagg Creek Member, a fluvial deposit of sandstone and conglomerate. Weems originally described this layer as lacking fossils, although later studies argued otherwise.Overlying the Stagg Creek Member was what Weems called the Falling Creek Member. As sediments began to slow down the rivers of the Stagg Creek Member, lacustrine deposits began to form. Rivers still managed to flow into the now dominant lakes from different directions, depositing a diverse assortment of sediments. This allowed the Falling Creek Member to contain a variety of rock types, including sandstone, shale, siltstone, and occasionally even coal. In addition, it is the most fossiliferous part of the Taylorsville basin, containing the fossils of not only fish and invertebrates, but also reptiles such as the heavily armored Doswellia.
The youngest and most geographically extensive member was the Newfound Member, named after the Newfound river. Outcrops belonging to this member either contain coarse sandstone and conglomerate, or much finer sandstone and siltstone. This member is believed to have been formed by a vast alluvial fan or delta created by southeastern-flowing rivers, and Araucarioxylon is occasionally found within the member.
Cornet & Olsen (1990)'s interpretation
In 1990, paleontologists Bruce Cornet & Paul Olsen described the Triassic fauna and flora of Virginia in detail. They argued that the Stagg Creek and Falling Creek Members actually coexisted at the same time, based on palynological correlations. In addition, they noted that the Stagg Creek actually preserved a few fossils, mainly of crustaceans and the abundant fish Dictyopyge.LeTourneau (2003)'s interpretation
In 2003, Columbia University geologist Peter LeTourneau became the first geologist to argue that the Doswell Formation was not the only Triassic geological layer in the Taylorsville Basin. He recognized the existence of a younger layer, which he called the King George Group. The King George Group was an extensive geological interval dominated by sandstone and conglomerate. LeTourneau also elevated the ranking of the Doswell Formation, renaming it to the Doswell Group. In conjunction with this, he also elevated the Falling Creek, and Newfound Members to formations. However, he also placed the Newfound Formation outside of the Doswell Group, instead placing it as the oldest unit of the King George Group.LeTourneau also evaluated Cornet & Olsen's claim that the Stagg Creek member coexisted with the Falling Creek Formation. He found that there were actually several distinct units grouped as part of the Stagg Creek unit. The original Stagg creek site which Weems based the member off of was found to belong to the middle portion of the Falling Creek Formation, therefore making the Stagg Creek Member part of that Formation. The portion of the Falling Creek Formation which was younger than the Stagg Creek Member was designated the Poor Farm Member, while the older portion was designated the Deer Creek Member. LeTourneau also added an additional formation to the Doswell Group, the South Anna Formation. This formation, the oldest section in the group, was very similar to the Stagg Creek Member in terms of its geological appearance and thickness, and as such South Anna outcroppings were originally considered to belong to the Stagg Creek Member according to Weems. LeTournea differentiated the two based on their age, as determined by palynological dating.
Weems, Tanner, & Lucas (2016)'s interpretation
In 2016, several paleontologists and geologists cooperated in a project which meant to correlate the individual Triassic basins of the Newark Supergroup with each other. They found that the Richmond and Taylorsville basins were likely deposited at the same time, a suspicion voiced earlier by Cornet & Olsen. In light of this revelation, they set out to link the formations and members of the Richmond Basin with those of the Taylorsville. They deranked the Doswell Group back to the Doswell Formation, and also found that the differences between the South Anna Formation and the Stagg Creek Member were not statistically significant. As a result, they abandoned the designation of the South Anna Formation, synonymizing it with the Stagg Creek Member as originally considered in 1980. They additionally found that the "barren beds" of the Richmond Basin were also synonymous with the Stagg Creek Member.The Falling Creek Formation was more thoroughly deconstructed. Weems, Tanner, & Lucas found that this formation was basically identical to the Vinita Member of the Richmond basin. As the Vinita member was named approximately 70 years earlier than the Falling Creek Formation, it was considered to take priority in naming. The Poor Farm and Deer Creek members of the Falling Creek Formation were also abandoned due to being poorly defined by LeTourneau.
These authors not only found the Doswell Formation to extend to the Richmond basin, but also to several other Triassic basins in Eastern Pennsylvania. For example, the "Irishtown beds" at the base of the Gettysburg basin were found to be a young layer of the Doswell Formation due to conchostracan dating. Lastly, an unusually old section of the Stockton Formation was also found to be a young part of the Doswell Formation due to preserving fossils of Calamops, a temnospondyl amphibian which lived at the same time as the Doswell Formation.
Paleobiota
Tetrapods
| Genus | Species | Sub-unit | Basin | Abundance | Notes | Images |
| Boreogomphodon | B. jeffersoni | Vinita member | Richmond | "hundreds of specimens" | A traversodontid cynodont | |
| Calamops | C. paludosis | Lahaska Creek member | Newark | A temnospondyl | ||
| Doswellia | D. kaltenbachi | Vinita member | Taylorsville, Richmond? | Partial skeleton, isolated bones | An archosauriform | |
| Euscolosuchus | E. olseni | Vinita member | Richmond | Armor scutes, vertebrae, and ribs | An archosauriform, possibly related to Crocodyliformes | |
| Gomphiosauridion | G. baileyae | Vinita member | Richmond | skull fragments | A procolophonid | |
| Lacertilia indet.? | Vinita member | Richmond | ||||
| Lissamphibia indet.? | Vinita member | Richmond | jaw fagment | |||
| Microconodon | M. tenuirostris | Vinita member | Richmond | Teeth and jaw bones | A cynodont | |
| Phytosauria indet. | Vinita member | Richmond, Taylorsville | teeth | |||
| Rauisuchidae indet. | Vinita member | Taylorsville? | limb bones | |||
| Rhynchocephalia indet. | Vinita member | Richmond | skull fagments | |||
| Uatchitodon | U. kroehleri | Vinita member | Richmond | teeth | A venomous reptile | |
| Xenodiphyodon | X. petraios | Vinita member | Richmond | partial jaw with teeth | A possible procolophonian |
Fish
Invertebrates
| Genus | Species | Sub-unit | Basin | Abundance | Notes | Images |
| Darwinula | Vinita member, Stagg Creek member | Richmond, Taylorsville | Ostracods | |||
| Gastropoda indet. | Vinita member | Richmond, Taylorsville | Snails | |||
| Laxitextella | L. multireticulata | Stagg Creek member, Vinita member, Irishtown member | Richmond, Taylorsville, Gettysburg | Abundant | A conchostracan | |
| Unionidae indet. | Vinita member | Taylorsville | Clams |
Pteridophytes (spore-bearing plants)
| Genus | Species | Sub-unit | Basin | Abundance | Notes | Images |
| Auriculophora | A. acrostichoides | Vinita member | Richmond | A marattialean fern | ||
| Cladophlebis | C. auriculata, C. mexicana | Vinita member | Richmond | A filicalean fern | ||
| Clathropteris | C. meniscoides | Vinita member | Richmond | A dipterid fern | ||
| Cyathocaulis | C. carolinensis | Vinita member | Richmond | A cyatheaceaen tree fern | ||
| Cyathoforma | C. carolinensis, C. minuta, C. penticarpa | Stagg Creek member, Vinita member | Richmond, Taylorsville | common | A large tree fern | |
| Danaeopsis | D. virginiensis | Vinita member | Richmond | A marattialean fern | ||
| Dicranopteris | D. sp. | Vinita member | Richmond | A forked fern | ||
| Equisetites | E. richmondensis, E. rogersii | Stagg Creek member, Vinita member | Richmond, Taylorsville | common | A horsetail | |
| Gleichenites | G. distans | Vinita member | Richmond | common | A forked fern | |
| Isoetodendron | I. striata | Vinita member | Richmond | A quillwort | ||
| Leptocyclotes | L. americana | Vinita member | Richmond | A quillwort | ||
| Lonchopteris | L. oblonga | Vinita member | Richmond | A fern | ||
| Mertensides | M. bullatus | Vinita member | Richmond | common | A fern | |
| Neocalamites | N. virginiensis,N. delawarensis | Stagg Creek member, Vinita member | Richmond, Taylorsville | common | A horsetail | |
| Osmundites | O. winterpockensis | Vinita member | Richmond | An osmunadcean fern | ||
| Pecopteris | P. rarinervis | Vinita member | Richmond | A type of leaf referable to several types of plants | ||
| Phlebopteris | P. smithii | Stagg Creek member | Taylorsville | A matoniacean fern | ||
| Pteridocaulis | P. rhombiformis, P. facialis | Vinita member | Richmond | Stem of a tree fern | ||
| Todites | T. linnaeaefolius | Stagg Creek member, Vinita member | Richmond, Taylorsville | common | A fern |
Gymnosperms
| Genus | Species | Sub-unit | Basin | Abundance | Notes | Images |
| Macrotaniopteris | M. crassinervis, M. magnifolia | Stagg Creek member, Vinita member | Richmond, Taylorsville | common | A cycad | |
| Plicarizamites | P. lanceolatus | Vinita member | Richmond | |||
| Podozamites | P. lanceolatus, P. tenuistriatus | Stagg Creek member, Vinita member | Richmond, Taylorsville | A type of leaf referable to conifers | ||
| Primaraucaria | P. wielandii | Vinita member | Richmond | fairly common | An araucariacean conifer | |
| Pterophyllum | P. affinae, P. braunianum, P. inaequale, P. giganteum, P. grandifolium, P. taxinum, P. tenuinervis | Stagg Creek member, Vinita member | Richmond, Taylorsville | Very common | A cycadeoid bennetitalean | |
| Sagenopteris | S. rhoifolia | Vinita member | Richmond | A seed fern | ||
| Sphenobaiera | S. striata | Stagg Creek member, Vinita member | Richmond, Taylorsville | common | A ginkgophyte | |
| Sphenopteris | S. sitholeyi | Stagg Creek member, Vinita member | Richmond, Taylorsville | common | A seed fern | |
| Sphenozamites | S. rogersianus | Stagg Creek member, Vinita member | Richmond, Taylorsville | common | A cycad | |
| Stangerites | S. obliqua, S. planus | Vinita member | Richmond | A cycad | ||
| Taeniopteris | T. diminuta | Vinita member | Richmond | common | Leaves of a Pentoxylon-like plant | |
| Triassiflorites | T. grandiflora | Vinita member | Richmond | A cone, possibly from a bennetitalean | ||
| Zamiostrobus | Z. lissocardius | Vinita member | Richmond | A cone from a cycad or conifer | ||
| Zamites | Z. powellii | Stagg Creek member | Taylorsville | A cycadeoid bennetitalean |