Fissurellidae


Fissurellidae, common name the keyhole limpets and slit limpets, is a taxonomic family of limpet-like sea snails, marine gastropod molluscs in the clade Vetigastropoda.
Their common name derives from the small hole in the apex of their cone-like shells. Although superficially resembling "true" limpets, they are in fact not closely related to them.

Distribution

The distribution of fissurellids is worldwide, from cold waters to tropical waters.

Habitat

Fissurellids live in habitats on and under rocks in the lower intertidal zones to deeper waters.

Shell description

Keyhole limpets somewhat resemble true limpets because of the simple conical shape of their shells, but in reality they are not closely related to true limpets, which are in the clade Patellogastropoda. This conical shape of the shell allows keyhole limpets to withstand wave attack on exposed rocks. The shell has a reticulate sculpture with strong radial ribs and lacks an operculum. The shell ranges from 3 mm to 13.2 cm. The great keyhole limpet measures up to 13.2 cm.
For respiration, the shells of fissurellids have a single apical or subapical perforation. This opening at the top allows a direct exit of exhalant water currents together with waste products from the mantle cavity. The water enters under the edge of the shell near the head and passes over large paired gills. Most young species in this family have a marginal slit in the middle of the anterior end of the spiral shell. Some species possess just a short internal groove at the anterior end. The paired organs in the mantle cavity represent a primitive condition in gastropods.
The soft body consists of a well-developed head, a short muzzle. It has a broad and flat foot and a well-developed mantle. This foot exerts a strong suction, adhering the keyhole limpet to its hard substratum. The mantle extends in some species partly or completely over the shell. The tentacles at the epipodium are well developed. The species in Medusafissurella have numerous subequal tentacles at the propodium, while the species in Dendrofissurella'' have an outgrowth with main trunk and side branches at the propodium. The eyes are situated on rudimentary pedicels at their outer bases. The sides are ornamented with short cirri. There are two, symmetrical branchial plumes. The anal siphon occupies the anterior notch or perforated summit of the shell.
In addition to the possession of this hole, slit or groove, keyhole limpets differ in several other ways both internally and externally from true limpets.

Feeding habits

Keyhole limpets are in essence herbivorous, feeding primarily on algae, but are also detritus feeders. A few species in the genera Diodora and Emarginella are carnivorous, feeding on sponges. Puncturella has been reported to digest diatoms and detritus. Puncturella aethiopica feeds mainly on Foraminifera.

Taxonomy

Family Fissurellidae Fleming, 1822
This classification was based by Bouchet & Rocroi on the studies by Keen , Christiaens and McMean.
Aktipis S.W., Boehm E. & Giribet G. then raised the tribe Diodorini to the status of subfamily Diodorinae.

Genera

Genera within the family Fissurellidae include:
; Genera brought into synonymy: