Gasteria
Gasteria is a genus of succulent plants, native to South Africa.
Naming
The genus is named for its stomach-shaped flowers. Common names include ox-tongue, cow-tongue, lawyer's tongue and, occasionally, mother-in-law's tongue.Description
Gasterias are recognisable from their thick, hard, succulent "tongue-shaped" leaves. Their inflorescence is also unique, with their curved, stomach-shaped flowers, which hang from inclined racemes.Distribution
The species of this genus are mostly native to the Eastern Cape Province, South Africa, where the bulk of the species occurs – especially in the small area between Grahamstown and Uniondale which enjoys rainfall throughout the year. However distribution of several species extends widely across the low-altitude coastal regions of the country, in an arched horseshoe shape across South Africa. At the one end of the genus's distribution, a species Gasteria pillansii extends into the far south-west corner of Namibia. At the other end, a species reaches the Lebombo mountains of Swaziland.Taxonomy
Gasteria is part of the family Asphodelaceae, subfamily Asphodeloideae. Closely related genera include Aloe and Haworthia, and the species of these genera are known to hybridise relatively easily with each other.Dividing Gasteria into species is extremely difficult, as each plant can be highly variable. One plant will look different depending on its location, its soil and its age. Young Gasteria plants typically look entirely different to older specimens.
In addition, the species tend to flow into each other in gradual transitions, with many intermediate forms, rather than being cleanly divided into discrete and separate species. Lastly, hybrids occur easily and naturally, whenever the range of two species overlap in habitat.
There is therefore considerable disagreement on how many species exist, with as many as 100 names being listed. Current studies tend to agree that there are between 16 and 23 species.
Taxonomy according to flower morphology
Using morphology, a traditional and widely accepted taxonomy was described in 1994, dividing the genus into 2 sections, 4 series, and 21 species.- Section Longiflorae
- *Series Longifoliae :
- **Gasteria batesiana G.D.Rowley – Limpopo, Mpumalanga, KwaZulu-Natal
- ***G. batesiana var. batesiana
- ***G. batesiana var. dolomitica – Mpumalanga
- **Gasteria barbae van Jaarsv. – southern coast of Cape Province, between Knysna and Plettenberg Bay
- **Gasteria tukhelensis van Jaarsv. – KwaZulu-Natal
- **Gasteria croucheri Baker – south-east Cape Province to east KwaZulu-Natal
- ***G. croucheri subsp. croucheri – south-east Cape Province to East KwaZulu-Natal
- ***G. croucheri subsp. pendulifolia Zonn. – KwaZulu-Natal
- ***G. croucheri subsp. pondoensis N.R.Crouch – south-east Cape Province
- **Gasteria acinacifolia Haw. – southern coast of Cape Province
- **Gasteria polita van Jaarsv. – Cape Province
- *Series Multifariae :
- **Gasteria excelsa Baker – Transkei, Eastern Cape
- **Gasteria pulchra Haw. – Cape Province
- **Gasteria carinata Duval – south-south-west Cape Province
- ***G. carinata var. carinata – Langeberg in Cape Province
- ***G. carinata var. retusa van Jaarsv. – Worcester & Heidelberg in Cape Province
- ***G. carinata var. thunbergii van Jaarsv.=G. thunbergii N.E.Br. – near Herbertsdale in Cape Province
- ***G. carinata var. verrucosa van Jaarsv. – Bredasdorp in Cape Province
- **Gasteria nitida Haw. – southern Cape Province
- ***G. nitida var. nitida – southern Cape Province
- ***G. nitida var. armstrongii van Jaarsv. / G. armstrongii Schönland – southern Cape Province
- **Gasteria ellaphieae van Jaarsv. – Kouga Dam in Cape Province
- **Gasteria glauca van Jaarsv – Cape Province
- **Gasteria vlokii van Jaarsv. – Great Swartberg + Witteberg in Cape Province
- Section Gasteria
- *Series Gasteria :
- **Gasteria obliqua Duval – south-east Cape Province; naturalized in Mexico
- ***=G. bicolor Duval
- ***=G. bicolor var. fallax van Jaarsv.
- ***=G. bicolor var. liliputana van Jaarsv.
- **Gasteria disticha Haw. – Robertson Karoo and surrounds, Western Cape
- ***G. disticha var. disticha – south-west and south-central Cape Province
- ***G. disticha var. langebergensis van Jaarsv. – western Cape Province
- ***G. disticha var. robusta van Jaarsv. – western Cape Province
- **Gasteria brachyphylla van Jaarsv. – Little Karoo in Cape Province
- ***G. brachyphylla var. bayeri van Jaarsv. – Little Karoo in Cape Province
- ***G. brachyphylla var. brachyphylla
- **Gasteria rawlinsonii Oberm. – Baviaanskloof in Cape Province
- **Gasteria glomerata van Jaarsv. – Kouga Dam in Cape Province
- **Gasteria baylissiana Rauh – Suurberg Range in Cape Province
- **Gasteria doreeniae van Jaarsv. & A.E.van Wyk – Cape Province
- *Series Namaquana :
- **Gasteria pillansii Kensit – Namibia, Cape Province
- ***G. pillansii var. var. ernesti-ruschii van Jaarsv. – south-west Namibia to north-west Cape Province
- ***G. pillansii var. hallii van Jaarsv. – western Cape Province
- ***G. pillansii var. pillansii'' – western Cape Province
Taxonomy according to genome
- Group A
- Gasteria rawlinsonii
- Group B - 8 rare and restricted inland species :
- *Gasteria armstrongii
- *Gasteria polita
- *Gasteria glomerata
- *Gasteria pulchra
- *Gasteria ellaphieae
- *Gasteria vlokii
- *Gasteria glauca
- *Gasteria nitida
- Group C - 5 widespread, distichous species, mainly from the west of southern Africa:
- *Gasteria brachyphylla
- *Gasteria bicolor
- *Gasteria disticha
- *Gasteria baylissiana
- *Gasteria pillansii
- Group D - 5 widespread, rosette-forming species, mainly large coastal species:
- *Gasteria excelsa
- *Gasteria croucheri
- *Gasteria acinacifolia
- *Gasteria carinata
- Group E
- *Gasteria batesiana
Gallery for identification
Western distichous group
Species with distichous, strap-shaped leaves which are usually without keels.Rare inland species
Large coastal group
Species generally form rosettes, with leaves usually bearing marginiform keels.Cultivation
Gasteria species are grown in well-drained, sandy soils in light shade. The species can all be propagated by off-sets and cuttings. They are also commonly propagated by seed. Germination usually occurs within 8 days but may take as long as one month depending on the species.Flowering times vary between species, but is usually in the spring & summer.
Those in the summer rainfall areas to the east, tend to always flower in spring to summer such as Gasteria batesiana, Gasteria croucheri & Gasteria acinacifolia.
Those in the areas which receive rainfall all year, usually flower also in later summer such as Gasteria excelsa, Gasteria nitida, Gasteria vlokii and Gasteria brachyphylla var. bayeri. Others in this region flower all year, but with a peak in the spring, such as Gasteria rawlinsonii, Gasteria bicolor & Gasteria carinata.
The western-most species vary in their flowering times, within the species. Gasteria pillansii in the far west, flowers in summer, except for its northern-most variety "var. ernesti-ruschii" which flowers in autumn.
Gasteria disticha usually flowers in spring, but in the far north of its range near Beaufort West it flowers in December.
Gasteria species are prone to Fusarium root rot, if they are over-watered.
The cultivar 'Little Warty' is a recipient of the Royal Horticultural Society's Award of Garden Merit.
Several hybrids with species in other related genera have been created in cultivation, such as between Gasteria and Aloe, and between Gasteria and Haworthia.