Gerenuk


The gerenuk, also known as the giraffe gazelle, is a long-necked antelope found in the Horn of Africa and the drier parts of East Africa. The sole member of the genus Litocranius, the gerenuk was first described by the naturalist Victor Brooke in 1879. It is characterised by its long, slender neck and limbs. The antelope is tall, and weighs between. Two types of colouration are clearly visible on the smooth coat: the reddish brown back or the "saddle", and the lighter flanks, fawn to buff. The horns, present only on males, are lyre-shaped. Curving backward then slightly forward, these measure.

Taxonomy and phylogeny

The gerenuk was first described by Victor Brooke in 1879 on the basis of three male specimens procured on "the mainland of Africa, north of the island of Zanzibar". Brooke used the scientific name Gazella walleri, on the request of Gerald Waller to name it after his deceased brother. The type locality was later corrected by John Kirk, who originally obtained the specimens on the "coast near the River Juba in southern Somaliland" before giving them to Waller. In 1886, Franz Friedrich Kohl proposed a new genus for the gerenuk, Litocranius. The common name derives from the Somali name for the animal ; the first recorded use of the name dates back to 1895. It is also known as the "giraffe gazelle" due to its similarity to the giraffe.
Two subspecies have been proposed, but these are considered to be independent species by some authors.
In 1997 Colin Groves proposed that Litocranius is a sister taxon of the similarly long-necked dibatag, but withdrew from this in 2000. A 1999 phylogenetic study based on cytochrome b and cytochrome c oxidase subunit III analysis showed that the tribe Antilopini, to which the gerenuk belongs, is monophyletic. In 2013, Eva Verena Bärmann and colleagues revised the phylogeny of tribe on the basis of nuclear and mitochondrial gene analysis. The cladogram prepared by them showed that the springbok forms a clade with the gerenuk; this clade is sister to the saiga and the genera Antilope, Eudorcas, Gazella and Nanger.

Description

The gerenuk is a notably tall, slender antelope that resembles gazelles. It is characterised by its long, slender neck and limbs, the flat, wedge-like head and the large, round eyes. Males are nearly tall, and the shorter females ; the head-and-body length is typically between. Males weigh between ; females are lighter, weighing. The species is sexually dimorphic. The tail, that ends in a black tuft, measures.
Two types of colouration are clearly visible on the smooth coat: the reddish brown dorsal parts, and the lighter flanks, fawn to buff. The underbelly and insides of the legs are cream in colour. The eyes and the mouth are surrounded by white fur. Females have a dark patch on the crown. The horns, present only on males, are lyre-like. Curving backward then slightly forward, these measure.
The gerenuk resembles the dibatag, with which it is sympatric in eastern and central Somalia and southeastern Ethiopia. Both are brachyodonts and share several facial and cranial features, along with a two-tone colouration of the coat and strong thick horns. However, there are also some features distinguishing it from the gerenuk, including major morphological differences in horns, horn cores, tail, postorbital area and basioccipital processes. The gerenuk has a longer, heavier neck and a shorter tail. A finer point of difference is the absence of an inward-curving lobe in the lower edge of the ear in the gerenuk. The subspecies of the gerenuk are similar in colouration; the southern gerenuk is the smaller of the two.

Ecology and behaviour

The gerenuk is a diurnal animal, though it typically stands or rests in shade during the noon. Foraging and feeding is the major activity throughout the day; females appear to spend longer time in feeding. The gerenuk may expose itself to rain, probably to cool its body. The social structure consists of small herds of two to six members. Herds typically comprise members of a single sex, though female herds additionally have juveniles. Some males lead a solitary life.
Fighting and travel are uncommon, possibly as a strategy to save energy for foraging. Both sexes maintain home ranges large, and might overlap. Those of males are scent-marked with preorbital gland secretions and guarded - hence these may be termed territories. The sedentary tendency of the antelope appears to increase with age.

Diet

Primarily a browser, the gerenuk feed on foliage of bushes as well as trees, shoots, herbs, flowers and fruits. It can reach higher branches and twigs better than other gazelles and antelopes by standing erect on its hindlegs and elongating its neck; this helps it reach over above the ground. Acacia species are eaten whenever available, while evergreen vegetation forms the diet during droughts. The pointed mouth assists in extracting leaves from thorny vegetation. The gerenuk does not drink water regularly. Major predators of the antelope include Cape hunting dogs, cheetahs, hyenas, lions and leopards.

Reproduction

Gerenuk reproduce throughout the year. Females reach sexual maturity at around one year, and males reach sexual maturity at 1.5 years, although in the wild they may only be successful after acquiring a territory. The gestation period is about seven months. They are born one at a time, weighing about at birth. Offspring were produced through artificial insemination for the first time in 2010 at White Oak Conservation in Yulee, Florida. Four female calves were born, and one of the four was later inseminated successfully by White Oak and SEZARC, creating a second generation of calves born from artificial insemination. Gerenuk can live thirteen years or more in captivity, and at least eight years in the wild.