List of non-marine molluscs of Great Britain
This list comprises 231 species of non-marine molluscs that have been recorded in the scientific literature as part of the fauna of the island of Great Britain; this total excludes species found only in hothouses and aquaria. The list includes terrestrial and aquatic gastropods, and aquatic bivalves. Molluscs that are fully marine are not included here, except for two marine pulmonate snails. In other words, this list includes land snails and slugs, and freshwater and brackish water snails. It also includes freshwater mussels and clams, including some that can tolerate brackish water.
Great Britain is a European island in the northeastern Atlantic, comprising the contiguous countries of England, Scotland and Wales. The mollusc fauna of the island of Ireland, which includes both the Republic of Ireland and Northern Ireland, is listed in another article: List of non-marine molluscs of Ireland.
A number of species of snails listed here are sometimes also found on lists of marine species. Two listed here, in the genera Onchidella and Otina, are fully marine in habitat, but are pulmonate snails that breathe air at low tide rather than having gills like most marine species. Other species listed here live in habitats that are intermediate between land and saltwater, or in brackish water habitats intermediate between freshwater and full-salinity saltwater.
Additional species are still being added to the list of the non-marine malacofauna of Great Britain. Four of the more recent discoveries are: Papillifera papillaris, first recorded in 1993 but probably the result of introductions with Italian statuary a century or more earlier; Selenochlamys ysbryda, a species new to science, which was first found in 2004; Candidula olisippensis, discovered on a Cornish cliff in 2011; and Monacha ocellata, found near Tilbury docks in 2017. Other species have been added to the list as a result of taxonomic revisions; for instance, only in 2009 was it recognised that snails previously known as Pupilla muscorum constituted two sibling species occupying different habitats, with both Pupilla muscorum sensu stricto and Pupilla pratensis occurring in Britain. A major revision of the slug fauna published in 2014, partly based on genetic sequencing, established that there were 20% more species than had previously been recognised. Not all these species have been definitely identified and some are formally undescribed.
In addition to the species that survive outdoors in Great Britain, there are also another 14 exotic gastropod species which live as "uninvited guests" in greenhouses and their enclosed aquaria. These species are known as "hothouse aliens", and are listed separately at the end of the main list. In this list these species are not counted as part of the fauna. Also not included are species such as Eobania vermiculata and Milax nigricans that have been found outdoors on single occasions but seem never to have established persisting populations.
The following table shows a summary of species numbers.
Non-marine molluscs of Great Britain | |
Gastropods land | 152 |
Gastropods aquatic | 48 |
Gastropods total: | 200 |
Bivalves freshwater | 31 |
Mollusc total: | 231 |
Gastropods introduced : | 31 land + 4 aquatic |
Bivalves introduced : | 3 |
Molluscs introduced in natural habitats, total: | 38–54 |
Gastropods living as "hothouse aliens" | 14 |
Bivalves living as "hothouse aliens" | 0 |
Conservation
Those species that are recognized as endangered are shown with an E after their name, see List of endangered species in the British Isles.Some species are protected in the United Kingdom under the Wildlife and Countryside Act 1981:
- Catinella arenaria and Myxas glutinosa are fully protected since 1981
- Margaritifera margaritifera is partly protected since 1991 and fully protected since 1998
- Helix pomatia is partly protected since 2008
- Monacha cartusiana was fully protected from 1981 to 1988.
Systematic list
The list is arranged by presumed biological affinity, rather than being alphabetical by family.A number of species are listed with subspecies, in cases where there are well-recognized subspecies in different parts of Europe. For some species a synonym is given, where the species may perhaps be better known under another name.
An attempt has been made to label the families as aquatic, terrestrial or intermediate, and an indication is given where it is thought that the species is introduced. Some introductions to Great Britain are quite ancient, dating from Roman times or even earlier, for example, Paleolithic. Those species that do not have a shell usually do not leave an archeological or fossil record, and therefore it is not always possible to determine whether they are native or introduced. Species are considered to be native, unless otherwise indicated on the list; this information is taken from Kerney, 1999, updated in the case of slugs with the opinions of Rowson et al. 2014.
Note: the images used to illustrate the list are mostly of specimens that were found in other countries.
[Gastropoda]
; Neritidae - aquatic- Theodoxus fluviatilis fluviatilis
- Acicula fusca
- Viviparus contectus
- Viviparus viviparus
- Assiminea grayana Fleming, 1828 - native?
- Paludinella littorina
- Marstoniopsis insubrica - synonym: Marstoniopsis scholtzi - native?
; Truncatellidae - terrestrial
; Valvatidae - aquatic
- Valvata cristata O. F. Müller, 1774
- Valvata macrostoma Mörch, 1864
- Valvata piscinalis
- Pomatias elegans
; Ellobiidae - terrestrial
- Carychium minimum Müller, 1774
- Carychium tridentatum
- Leucophytia bidentata
- Myosotella denticulata
- Myosotella myosotis
; Onchidiidae - aquatic
; Lymnaeidae - aquatic
- Galba truncatula
- Omphiscola glabra
- Lymnaea palustris = Stagnicola palustris
- Lymnaea stagnalis
- Lymnaea fusca
- Radix auricularia
- Radix balthica
- Radix peregra
- Myxas glutinosa E
; Planorbidae - aquatic
; Succineidae - terrestrial
- Quickella arenaria = Catinella arenaria
- Succinella oblonga Draparnaud, 1801
- Succinea putris Linnaeus, 1758
- Oxyloma elegans elegans = Oxyloma pfeifferi
- Oxyloma sarsii
- Azeca goodalli
- Cochlicopa cf. lubrica = Cochlicopa lubrica
- Cochlicopa cf. lubricella = Cochlicopa lubricella
- Pyramidula pusilla
- Columella edentula
- Columella aspera Walden, 1966
- Truncatellina cylindrica
- Truncatellina callicratis - probably native
- Vertigo pusilla O. F. Müller, 1774
- Vertigo alpestris Alder, 1838
- Vertigo antivertigo
- Vertigo genesii
- Vertigo geyeri Lindholm, 1925
- Vertigo lilljeborgi
- Vertigo modesta arctica
- Vertigo moulinsiana
- Vertigo pusilla O. F. Müller, 1774
- Vertigo pygmaea
- Vertigo substriata
- Vertigo angustior Jeffreys, 1830
; Chondrinidae - terrestrial
; Valloniidae - terrestrial
- Vallonia costata
- Vallonia pulchella
- Vallonia cf. excentrica = Vallonia excentrica Sterke, 1892
- Acanthinula aculeata
- Spermodea lamellata
; Enidae - terrestrial
; Arionidae - terrestrial
; Euconulidae - terrestrial
- Euconulus cf. fulvus ?= Euconulus fulvus
- Euconulus cf. alderi ?= Euconulus alderi
- Zonitoides nitidus
- Zonitoides excavatus
; Oxychilidae - terrestrial
; Limacidae - terrestrial
; Ferussaciidae - terrestrial
- Cecilioides acicula - probably introduced
- Alinda biplicata biplicata - introduced?
- Balea perversa
- Balea heydeni Von Maltzan, 1881
- Clausilia bidentata bidentata
- Clausilia dubia dubia Draparnaud, 1805 - probaby native
- Cochlodina laminata
- Macrogastra rolphii
- Papillifera papillaris - introduced
; Testacellidae - terrestrial
; Helicodontidae - terrestrial
- Helicodonta obvoluta obvoluta - probably native
- Cochlicella acuta - introduced?
- Cochlicella barbara - introduced
; Hygromiidae - terrestrial
- Ashfordia granulata = Monacha granulata
- Candidula gigaxii - introduced?
- Candidula intersecta - probably introduced
- Candidula olisippensis - introduced
- Cernuella aginnica - introduced
- Cernuella neglecta - introduced, now extinct in G.B.
- Cernuella virgata - probably introduced
- Helicella itala itala
- Hygromia cinctella - introduced
- Hygromia limbata limbata - introduced
- Monacha cantiana - introduced
- Monacha cartusiana - introduced?
- Monacha ocellata - introduced
- Ponentina subvirescens - probably native
- Pseudotrichia rubiginosa - probably native
- Trochoidea elegans - introduced
- Trochulus hispidus
- Trochulus sericeus
- Trochulus striolatus - indigenous?
- Zenobiella subrufescens
- Fruticicola fruticum - probably introduced, now extinct in G.B.
; Helicidae - terrestrial
[Bivalvia]
; Margaritiferidae - aquatic- Margaritifera margaritifera E
; Unionidae - aquatic
- Unio pictorum
- Unio tumidus Philipsson, 1788
- Anodonta anatina
- Anodonta cygnea
- Pseudanodonta complanata E
- Sphaerium corneum
- Sphaerium rivicola
- Sphaerium nucleus
- Sphaerium solidum
- Musculium lacustre
- Musculium transversum
- Pisidium amnicum
- Pisidium casertanum
- Pisidium conventus
- Pisidium henslowanum
- Pisidium hibernicum Westerlund, 1894
- Pisidium lilljeborgii
- Pisidium milium Held, 1836
- Pisidium moitessierianum Paladilhe, 1866
- Pisidium nitidum Jenyns, 1832
- Pisidium obtusale
- Pisidium personatum Malm, 1855
- Pisidium pulchellum
- Pisidium pseudosphaerium Favre, 1927
- Pisidium subtruncatum Malm, 1855
- Pisidium supinum A. Schmidt, 1851
- Pisidium tenuilineatum Stelfox, 1918 E
- Dreissena polymorpha - introduced
- Mytilopsis leucophaeata - introduced
- Corbicula fluminea - introduced
List of "hothouse alien" species
on orchid plants
; Thiaridae - aquatic
- Melanoides tuberculata
- Zonitoides arboreus
- Radix rubiginosa, native to Indo-China and Indonesia
- Gyraulus chinensis
- Planorbella duryi
- Pleurodiscus balmei