Littoraria Littoraria is a genus of sea snails , marine gastropod mollusks in the family Littorinidae , the winkles or periwinkles. There are more than fifty species in this genus of which more than 20 species are believed to be synonyms of Littoraria scabra , a very variable species. Many of the species in this genus occur in the Indo-West Pacific region and in the Tropical Eastern Pacific , where they are found in large numbers on the trunks, trees and prop roots of tropical mangrove forests a few metres above high tide level. These snails feed on the thin film of algae , epiphytes, fungi , diatoms and leaf epidermis of these mangroves. The species living on higher levels of the trees have thinner shells, and are more variable in shell colour. Within this genus, Littoraria aberrans is the only ovoviviparous species with an metamorphosis.Species Species in the genus Littoraria include:Littoraria aberrans Philippi , 1846Littoraria ahenea Reid, 1986Littoraria albicans Metcalfe, 1852Littoraria angulifera Lamarck , 1822 - mangrove periwinkle *Littoraria angulifera hessei Boettger 1912 Littoraria ardouiniana Littoraria articulata Philippi, 1846Littoraria bengalensis Reid, 2001Littoraria carinifera Menke , 1830Littoraria cingulata Philippi, 1846Littoraria cingulifera Dunker , 1845Littoraria coccinea Littoraria conica Philippi, 1846Littoraria delicatula Nevill , 1885Littoraria filosa Littoraria fasciata Sowerby , 1832† Littoraria flammea Philippi, 1847 Littoraria flava King & Broderip, 1832Littoraria flavaosa King & Broderip, 1832Littoraria ianthostoma Stuckey & D.G. Reid, 2002Littoraria intermedia Philippi, 1846Littoraria irrorata Say , 1822 - marsh periwinkle Littoraria kraussi Rosewater, 1970 Littoraria lutea Philippi, 1847Littoraria luteola Quoy & Gaimard , 1833 Littoraria mauritiana Lamarck, 1822Littoraria melanostoma Gray , 1839Littoraria miodelicatula Oyama, 1950Littoraria nebulosa Lamarck, 1822 - cloudy periwinkleLittoraria obesa Gmelin , 1791Littoraria pallescens Philippi, 1846Littoraria philippiana Reeve , 1857Littoraria pintado Wood, 1820 *Littoraria pintado pullata Carpenter , 1864 Littoraria planaxis Philippi, 1846Littoraria punctata Philippi, 1846Littoraria rosewateri Reid, 1999Littoraria scabra Philippi, 1847Littoraria sinensis Philippi, 1847Littoraria strigata Philippi, 1846Littoraria subvittata Reid, 1986Littoraria sulculosa Philippi, 1846Littoraria tessellata Philippi, 1847Littoraria undulata Gray, 1839Littoraria varia Sowerby, 1832Littoraria variegata Souleyet , in Eydoux & Souleyet, 1852Littoraria vespacea Reid, 1986Littoraria zebra Donovan , 1825 - zebra periwinkle ;Species brought into synonymy :Littoraria glabrata Philippi, 1846 : synonym of Littoraria coccinea glabrata
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