Memecylon


Memecylon is a plant group in Melastomataceae. It consists of 350-400 species of small to medium-sized trees and shrubs occurring in the Old World tropics. Memecylon is a monophyletic group basal to the Melastomataceae clade. Memecylon taxa have more than 600 published basionyms. Diversity of this group is concentrated in tropical Africa, Madagascar, Sri Lanka, India and Malaysia.

Etymology

The name Memecylon is derived from 'memaecylon' as used by ancient Greek philosophers Dioscorides and Pliny to describe the red fruits of Arbutus unedo, an unrelated plant group, alluding to the pink to reddish berries often produced by Memecylon. Some vernacular names in different regions of the world are given below.
English: Blue mist plant, Hindi: Anjan, Malayalam: Aattukanala , Oriya: Neymaru, Sinhala: Korakaha, Welikaha, Tamil: Kaya.

Morphology

Memecylon sensu lato can be diagnosed by exstipulate leaves, four-merous bisexual flowers, anthers opening by slits, enlarged connectives bearing terpenoid secreting glands and berries. Memecylon sensu stricto can be distinguished from other Memecyloids by obscure nervation on leaves, non-glandular roughened leaf surface having branched sclerids, imbricate calyx, unilocular ovary and large embryo with thick and convoluted cotyledons.

Stems

Species of larger stature have a characteristic brown bark with narrow and sharp furrows, most are small single stemmed trees. However, the bark of many species of smaller stature is varied and may be papery white or smooth dark red black.

Leaves

Leaves are opposite, short-stalked, elliptic or ovate, mostly with widely spaced pinnate nerves either visible or obscure. Leaves along the twig are all the same size, shiny, glabrous, with entire margins, the node has a characteristic scar between the leaves, the twig bark is typically red, striated and flaky.

Flowers

The inflorescence is typically dense and axillary. The florets are small compared to the other taxa in Melastomataceae, with short fleshy corolla parts. Cymes are bracteate, usually thyrsoid to umbel shaped, often condensed to sessile fascicles of flowers or a few-flowered heads at tips of peduncles. The florets are white or violet, the stamens blue or violet, usually obvious in aggregates, from axillary clusters. Flowers are bisexual, have inferior ovaries, but the parts are otherwise free. The calyx is valvate and there are twice as many stamens as petals.

Fruits

The fruit is from an inferior ovary, typically axillary. The calyx remnants are persistent, and are sometimes blue-black. Fruits are globose or occasionally ellipsoid, pulpy or juicy with one large seed.

Distribution

This group is distributed in approximately 53 countries of the Old World tropics and occupies a wide range of habitats. Memecylon taxa have been reported from montane forests, tropical lowland forests, grasslands, tropical rainforests with low to high rainfall, rocky mountain regions and regions with low to high temperature and a considerable overlap between ranges of different taxa. Most of the plants in this group are regionally or locally endemic. Endemic Memecylon are reported from 21 countries. The global distribution of this group is shown in the following Figure and some of the countries which have endemic Memecylon species are listed in the table.
geographic occurrence data and locality data in NCBI database
Table1. Number of Memecylon taxa reported from different countries
CountryTotal Number of Memecylon
Andamans9
Angola4
Australia3
Borneo18
Cambodia10
Cameroon27
Ceram Island3
Central African Republic2
China16
Congo10
Democratic Republic of Congo1
Equatorial Guinea7
Fiji3
Gabon24
Ghana4
Guinea6
India39
Indonesia14
Ivory Coast6
Java9
Kenya6
Laos7
Liberia9
Madagascar138
Malawi2
Malaysia29
Mascarene Islands4
Mauritius5
Mayotte1
Mozambique3
Myanmar12
New Guinea14
Nicobar Islands2
Nigeria5
Philippines16
Papua New Guinea9
Reunion2
Seychelle2
Sierra Leone4
Simalue Island2
Singapore12
South Africa2
Sri Lanka32
Sumatra10
Taiwan2
Tanzania10
Thailand21
Tonga1
Uganda1
Vanuatu1
Vietnam15
Zambia2
Zimbabwe1

Phylogeny

Phylogeny based on morphological treatments

The classification of Memecylon has switched back and forth among families Melastomataceae, Myrtaceae and Memecylaceae based on morphological, anatomical and molecular evidence. Early classifications excluded Memecylon and Mouriri from the Melastomataceae mainly based on placentation and seed characters. Subsequent treatments preferred to treat Memecylon as a member of Myrtaceae. Again, this group has been included in Melastomataceae by Cronquist. After that, morphological and anatomical character analysis of the Melastomataceae and their traditional allies by Renner identified two major lineages, and in that classification, Memecylon was placed in the Memecylaceae. Synapomorphies used in this phylogenetic analysis were anther connective and dehiscence, dorsal glands on stamen connectives, endothecium, placentation, locules, seeds, leaf venation, terminal leaf sclereids, paracytic stomata, stomata shape, leaf sclereids, indumentum, ant and mite domatia, wood and growth form characters, which excluded Memecylaceae from Melastomataceae. However, in Memecylon some characters such as seasonal flowering and small size of flowers contributed to the difficulty of assessing relationships based on the morphology. Later, several groups have been either included in broadly defined Memecylaceae or segregated from it. As a result, Memecylaceae comprised the groups Memecylon, Lijndenia, Mouriri, and :pt:Votomita|Votomita.

Phylogeny based on molecular treatments

Molecular phylogenetic analyses using rbcL sequence data showed that the Melastomataceae lineage consists of a subclade formed by Oliniaceae, Penaeaeae, Phynchocalycaceae and Alzateaceae sister to a subclade formed by Memecylaceae and Melastomataceae sensu stricto. Parsimony analysis showed distinct Myrtaceae and Melastomataceae clades. Subsequently, Bayesian analyses using chloroplast DNA sequences from the rbcL and ndhF genes, and parsimony and maximum likelihood analyses on rRNA, rbcL and atpB sequences, have shown that Memecylaceae is a sister group to Melastomataceae. Later studies using combined exon and intron sequences of the nuclear glyceraldehyde 3-phosphate dehydrogenase gene has supported the monophyly of Memecylon. However, the APG IV system of classification, a system of revised and updated classification of flowering plants, now recognizes the groups of Memecylaceae within a broader circumscription of Melastomataceae.
The most recent phylogenetic analysis of Memecylon included mostly African taxa and was based on two nuclear ribosomal regions, ETS and ITS. Memecylon species discovery in Madagascar increased, and comprehensive taxonomic revisions brought the total number up to 138 species, making Memecylon one of the most species-rich groups in the woody flora of Madagascar. In addition to this high species richness, a high degree of endemism of Memecylon indicated the importance of considering ecogeographic factors when making species identifications in this group. Therefore, in the latest molecular phylogeny, a combined study of ecogeographic factors has been taken into account.

Pollination and seed dispersal

The dense and axillary showy clusters of Memecylon florets do not produce nectar. These flowers are visited by pollen-gathering bees who vibrate or manipulate the anthers. Anthers open by longitudinal slits and exposed pollen invites pollen gathering bees. Anther appendages serve as a hold for bees’ legs. These flowers have terpenoid secreting glands and bees collect terpenoids. Therefore, buzz pollination is also favored. Berries are dispersed by birds and mammals. Populations of Memecylon are widely scattered within the forests as would be expected in bird-dispersed species.

Ecology

Memecylon produce flowers and fruits more regularly than many trees of the equatorial forests. It provides a food supply for wildlife as a source of fruits. Different sympatric groups appear to segregate mainly by soil moisture.

Ethnobotany

Memecylon has economic, medicinal and horticultural values. A yellow dye and a mordant can be extracted from the leaves. They are traditionally used for dyeing silk in Thailand and the robes of Buddhist monks in Sri Lanka. Fruits are edible and some are used as spices. This group provides hard and valuable timber used for building houses and boats. Wood is used to make rafters, house posts, fuel wood, charcoal, tools, and handles. An infusion of leaves is used as astringent for ophthalmia. Further, the leaves are used in the treatment of gonorrhea, or when mixed with several other ingredients, they make good formulations for external use. The bark is applied as a poultice to bruises. Root and heartwood decoctions are used to bring down fever associated with colds, chicken pox and measles.

A list of ''Memecylon'' species

  1. Memecylon aberrans H. Perrier
  3. Memecylon acrocarpum Bakh. f.
  5. Memecylon acrogenum R.D. Stone
  7. Memecylon acuminatissimum Blume
  9. Memecylon acuminatum Sm.
  11. Memecylon aequidianum Jacq.-Fél.
  13. Memecylon affine Merr.
  15. Memecylon afzelii G. Don
  17. Memecylon agastyamalaianum E.S.S. Kumar, Antony & A.E.S. Khan
  19. Memecylon alatum Aug. DC.
  21. Memecylon albescens Jacq.-Fél.
  23. Memecylon ambrense Jacq.-Fél.
  25. Memecylon amherstianum C.B. Clarke
  27. Memecylon amoenum Jacq.-Fél.
  29. Memecylon amplexicaule Roxb.
  31. Memecylon amplifolium R.D. Stone
  33. Memecylon amshoffiae Jacq.-Fél.
  35. Memecylon andamanicum King
  37. Memecylon angulatum Rchb. ex DC.
  39. Memecylon angustifolium Wight
  41. Memecylon antseranense Jacq.-Fél.
  43. Memecylon apoense Elmer
  45. Memecylon arcuatomarginatum Gilg ex Engl.
  47. Memecylon argenteum K. Bremer
  49. Memecylon arnhemense Whiffin
  50. Memecylon arnottianum Wight ex Thwaites
  52. Memecylon auratifolium H. Perrier
  54. Memecylon aylmeri Hutch. & Dalziel
  56. Memecylon azurinii Quisumb. & Merr.
  58. Memecylon bachmannii Engl.
  60. Memecylon bakerianum Hutch. & Dalziel; nom. illeg.
  62. Memecylon balakrishnanii Lakshmin. & S.P. Mathew
  64. Memecylon basilanense Merr.
  66. Memecylon batekeanum R.D. Stone & G.M. Walters
  67. Memecylon bequaertii De Wild.
  69. Memecylon bernieri Cogn.
  71. Memecylon boinense H. Perrier
  73. Memecylon brachybotrys Merr.
  75. Memecylon bracteatum Jacq.-Fél.
  77. Memecylon bracteolatum Bakh. f.
  79. Memecylon brahense Jacq.-Fél.
  81. Memecylon bremeri M.B. Viswan.
  83. Memecylon bretelerianum Jacq.-Fél.
  85. Memecylon buxifolium Blume
  87. Memecylon buxoides Wickens
  89. Memecylon caeruleum Jack
  91. Memecylon calderense A. Gray
  93. Memecylon calophyllum Gilg
  95. Memecylon calyptratum K. Bremer
  97. Memecylon campanulatum C.B. Clarke
  99. Memecylon candidum Gilg
  101. Memecylon candolleanum Cogn.
  103. Memecylon cantleyi Ridl.
  105. Memecylon capense Eckl. & Zeyh.
  107. Memecylon capitellatum sensu Span.
  109. Memecylon capuronii Jacq.-Fél.
  111. Memecylon cardiophyllum Cogn.
  113. Memecylon caudatum Craib
  115. Memecylon celastrinum Kurz
  117. Memecylon celebicum Bakh. f.
  119. Memecylon ceramense Bakh. f.
  121. Memecylon chevalieri Guillaumin
  123. Memecylon cinereum King
  125. Memecylon claessensii De Wild.
  127. Memecylon clarkeanum Cogn.
  129. Memecylon clausiflorum Naudin
  131. Memecylon clavistaminum Jacq.-Fél.
  133. Memecylon cogniauxii Gilg
  135. Memecylon confertiflorum Cogn.
  137. Memecylon confine Blume
  139. Memecylon confusum Blume
  141. Memecylon conocarpum Lauterb. & K. Schum.
  143. Memecylon constrictum Craib
  145. Memecylon cordatum Wall.; nom. nud.
  147. Memecylon cordifolium Merr.
  149. Memecylon corticosum Ridl.
  151. Memecylon corymbiforme H. Perrier
  153. Memecylon coursianum Jacq.-Fél.
  155. Memecylon crassifolium Bakh. f.
  157. Memecylon crassinerve Blume
  159. Memecylon crassipetiolum Jacq.-Fél.
  161. Memecylon cumingianum C. Presl
  163. Memecylon cumingii Naudin
  165. Memecylon cuneatum Thwaites
  167. Memecylon curtisii Burkill & M.R. Hend.
  169. Memecylon cyaneum De Wild.
  171. Memecylon cyanocarpum C.Y. Wu; nom. illeg.
  173. Memecylon dalleizettei H. Perrier
  175. Memecylon dallmannense Ohwi
  177. Memecylon dasyanthum Gilg & Lederm. ex Engl.
  179. Memecylon deccanense C.B. Clarke
  181. Memecylon delphinense H. Perrier
  183. Memecylon deminutum Brenan
  185. Memecylon densiflorum Merr.
  187. Memecylon dichotomum King
  189. Memecylon diluviorum Exell
  191. Memecylon dioicum Cogn.
  193. Memecylon discolor Cogn.
  195. Memecylon dissitum Craib
  197. Memecylon dolichophyllum Naudin
  199. Memecylon dubium Jacq.-Fél.
  201. Memecylon durum Cogn.
  203. Memecylon edule Roxb.
  205. Memecylon eduliforme' Aug. DC.
  207. Memecylon eglandulosum H. Perrier
  209. Memecylon elaeagni Blume
  211. Memecylon elegans Kurz
  213. Memecylon elegantulum Thwaites
  215. Memecylon ellipticum Thwaites
  217. Memecylon elliptifolium Merr.
  219. Memecylon elongatum Merr.
  221. Memecylon englerianum Cogn.
  223. Memecylon erythranthum Gilg
  225. Memecylon excelsum Blume
  227. Memecylon faucherei Danguy
  229. Memecylon fernandesiorum Jacq.-Fél.
  231. Memecylon fernandianum Gilg ex Engl.
  233. Memecylon ferreum Blume
  235. Memecylon fianarantse Jacq.-Fél.
  237. Memecylon flavescens Gamble
  239. Memecylon flavovirens Baker
  241. Memecylon fleuryi Jacq.-Fél.
  243. Memecylon floribundum Blume
  245. Memecylon floridum Ridl.
  247. Memecylon fragrans A. Fern. & R. Fern.
  249. Memecylon fruticosum King
  251. Memecylon fuscescens Thwaites
  253. Memecylon galeatum H. Perrier
  255. Memecylon garcinioides Blume
  256. Memecylon gardneri Thwaites
  257. Memecylon geddesianum Craib
  259. Memecylon geoffrayi Guillaumin
  261. Memecylon germainii A. Fern. & R. Fern.
  263. Memecylon gibbosum Bakh. f.
  265. Memecylon giganteum Alston
  267. Memecylon gigantifolium Elmer
  269. Memecylon gilgianum Exell
  271. Memecylon gitingense Elmer
  273. Memecylon gracile Bedd.
  275. Memecylon gracilipedicellatum Jacq.-Fél.
  277. Memecylon gracilipes Ridl.; nom. illeg.
  279. Memecylon gracillimum Alston
  281. Memecylon grande sensu Blume
  283. Memecylon greenwayi Brenan
  285. Memecylon griffithianum Naudin
  287. Memecylon hainanense Merr. & Chun
  289. Memecylon harmandii Guillaumin
  291. Memecylon helferi Cogn.
  293. Memecylon heyneanum sensu Wight
  295. Memecylon hookeri sensu A.N. Henry & Subr.
  297. Memecylon huillense A. Fern. & R. Fern.
  299. Memecylon hullettii King
  301. Memecylon humbertii H. Perrier
  303. Memecylon hylandii Whiffin
  304. Memecylon idukkianum Nampy & Radh
  306. Memecylon impressivenum R.D. Stone
  308. Memecylon inalatum Jacq.-Fél.
  310. Memecylon infuscatum Jacq.-Fél.
  312. Memecylon insigne Mansf.
  314. Memecylon insperatum A.C. Sm.
  316. Memecylon insulare A. Fern. & R. Fern.
  318. Memecylon interjectum R.D. Stone
  320. Memecylon isaloense Jacq.-Fél.
  322. Memecylon ivohibense Jacq.-Fél.
  324. Memecylon jadhavii K.N. Reddy, C.S. Reddy & V.S. Raju; nom. inval.
  326. Memecylon klaineanum Jacq.-Fél.
  328. Memecylon kollimalayana M.B. Viswan.
  330. Memecylon kratense Craib
  332. Memecylon kunstleri King
  334. Memecylon lancifolium Ridl.
  336. Memecylon langbianense Guillaumin
  338. Memecylon laruei Merr.
  340. Memecylon lateriflorum Bremek.
  342. Memecylon latifolium Bojer; nom. nud.
  344. Memecylon laurentii De Wild.
  346. Memecylon laureolum Jacq.-Fél.
  348. Memecylon laurinum Blume
  350. Memecylon lawsonii Gamble
  352. Memecylon laxiflorum Wall. ex Ridl.; nom. illeg.
  354. Memecylon leucanthum Thwaites
  356. Memecylon leucocarpum Gilg
  358. Memecylon liberiae Gilg ex Engl.
  360. Memecylon ligustrifolium Champ. ex Benth.
  362. Memecylon lijndenia Kuntze
  364. Memecylon lilacinum Zoll. & Moritzi
  366. Memecylon littorale Merr.
  368. Memecylon loheri Merr.
  370. Memecylon longicuspe Baker
  372. Memecylon longifolium Cogn.
  374. Memecylon longipetalum H. Perrier
  376. Memecylon louvelianum H. Perrier
  378. Memecylon luchuenense C. Chen
  380. Memecylon lucidum C. Presl
  382. Memecylon lureri Naudin
  384. Memecylon lushingtonii Gamble
  386. Memecylon macrocarpum Thwaites
  388. Memecylon macrodendron Gilg ex Engl.
  390. Memecylon macrophyllum Thwaites
  392. Memecylon madgolense Gamble
  394. Memecylon magnifoliatum A. Fern. & R. Fern.
  396. Memecylon malaccense Ridl.
  398. Memecylon mandrarense H. Perrier
  400. Memecylon mangiferoides Jacq.-Fél.
  402. Memecylon manickamii Murugan, Sundaresan & Jothi
  403. Memecylon mayottense R.D. Stone
  405. Memecylon megacarpum Furtado
  407. Memecylon megaspermum Jacq.-Fél.
  408. Memecylon memoratum Jacq.-Fél.
  410. Memecylon merguicum King
  412. Memecylon minimifolium H. Perrier
  414. Memecylon minutiflorum Miq.
  416. Memecylon mocquerysii Aug. DC.
  418. Memecylon monchyanum Backer; nom. inval.
  420. Memecylon mouririoides Jacq.-Fél.
  422. Memecylon multiflorum Bakh. f.
  424. Memecylon multinode Jacq.-Fél.
  426. Memecylon mundanthuraianum M.B.Viswan. & Manik.
  428. Memecylon myrianthum Gilg
  430. Memecylon myricoides Naudin
  432. Memecylon myrtiforme Naudin
  434. Memecylon myrtilloides Markgr.
  436. Memecylon natalense Markgr.
  438. Memecylon nigrescens Engl.; nom. illeg.
  440. Memecylon nodosum Gilg ex Engl.
  442. Memecylon normandii Jacq.-Fél.
  444. Memecylon novoguineense Baker f.
  446. Memecylon obscurinerve Merr.
  448. Memecylon obtusifolium Merr.
  450. Memecylon occultum Jacq.-Fél.
  452. Memecylon ochroleucum Bakh. f.
  454. Memecylon octocostatum Merr. & Chun
  456. Memecylon odoratum Elmer
  458. Memecylon oligophlebium Merr.
  460. Memecylon orbiculare Thwaites
  462. Memecylon oubanguianum Jacq.-Fél.
  464. Memecylon ovatifolium Wickens
  466. Memecylon ovatum Sm.
  468. Memecylon ovoideum Thwaites
  470. Memecylon pachyphyllum Merr.
  472. Memecylon palawanense Elmer
  474. Memecylon pallidum Merr.
  476. Memecylon paniculatum Jack
  478. Memecylon papuanum Merr. & L.M. Perry
  480. Memecylon paradoxum Jacq.-Fél.
  482. Memecylon parviflorum Blanco
  483. Memecylon parvifolium Thwaites
  485. Memecylon pauciflorum Blume
  487. Memecylon pedunculatum Jacq.-Fél.
  489. Memecylon pendulum Chih C. Wang, Y. H. Tseng, Y. T. Chen & Kun C. Chang
  491. Memecylon peracuminatum H. Perrier
  493. Memecylon perangustum Jacq.-Fél.
  495. Memecylon perditum R.D. Stone
  497. Memecylon pergamentaceum Cogn.
  499. Memecylon perrieri Danguy
  501. Memecylon pervilleanum Naudin
  503. Memecylon petiolatum Trimen ex Alston
  505. Memecylon phanerophlebium Merr.
  507. Memecylon phyllanthifolium Thwaites ex Bedd.
  509. Memecylon pierrei Hance
  511. Memecylon pileatum Jacq.-Fél.
  513. Memecylon planifolium Jacq.-Fél.
  515. Memecylon plebejum Kurz
  517. Memecylon polyanthemos Hook. f.
  519. Memecylon polyanthum H.L. Li
  521. Memecylon polyneuron Gilg
  523. Memecylon ponmudianum Sivu, N.S.Pradeep & Pandur.
  525. Memecylon preslianum Triana; nom. illeg.
  527. Memecylon procerum Thwaites
  529. Memecylon protrusum Bakh. f.
  531. Memecylon prunifolium Naudin
  533. Memecylon pseudomyrtiforme H. Perrier
  535. Memecylon pterocarpum H. Perrier
  537. Memecylon pterocaule Merr.
  539. Memecylon pterocladum R.D. Stone
  541. Memecylon pteropus Merr.
  543. Memecylon pubescens King
  544. Memecylon pulvinatum Jacq.-Fél.
  546. Memecylon punctatum C. Presl
  548. Memecylon purpurascens Jacq.-Fél.
  550. Memecylon pusilliflorum Cogn.
  552. Memecylon pyrifolium Naudin; nom. illeg.
  554. Memecylon ramosii Merr.
  556. Memecylon ramosum Jacq.-Fél.
  558. Memecylon randerianum S.M. Almeida & M.R. Almeida
  560. Memecylon rauschianum Gilg & Lederm. ex Engl.
  562. Memecylon revolutum Thwaites
  564. Memecylon reygaertii De Wild.
  566. Memecylon rhamnoideum Naudin
  568. Memecylon rhinophyllum Thwaites
  570. Memecylon rhodophyllum Bakh. f.
  572. Memecylon ridleyi Cogn. ex Ridl.; nom. nud.
  574. Memecylon rivulare K. Bremer
  576. Memecylon roseum H. Perrier
  578. Memecylon rostratum Thwaites
  580. Memecylon rotundatum Cogn.
  582. Memecylon royenii Blume
  584. Memecylon rubiflorum Jacq.-Fél.
  586. Memecylon rubrocaeruleum Thwaites
  588. Memecylon ruptile K. Bremer
  590. Memecylon sabulosum Jacq.-Fél.
  592. Memecylon salicifolium Jacq.-Fél.
  594. Memecylon sambiranense H. Perrier
  596. Memecylon sapinii De Wild.
  598. Memecylon schliebenii Markgr.
  600. Memecylon schraderbergense Mansf.
  602. Memecylon schumannianum Mansf.
  604. Memecylon scolopacinum Ridl.
  606. Memecylon scutellatum sensu Seem.
  608. Memecylon sejunctum R.D. Stone
  610. Memecylon semseii A. Fern. & R. Fern.
  612. Memecylon sepicanum Mansf.
  614. Memecylon sessile A. Chev.
  615. Memecylon sessilicarpum A. Fern. & R. Fern.
  617. Memecylon sessilifolium Merr.
  619. Memecylon simii Stapf
  621. Memecylon sisparense Gamble
  623. Memecylon sitanum Jacq.-Fél.
  625. Memecylon sivadasanii N. Mohanan, Ravi, Kiran Raj & Shaju
  627. Memecylon sorsogonense Elmer
  629. Memecylon sousae A. Fern. & R. Fern.
  631. Memecylon sparsiflorum Bojer; nom. nud.
  633. Memecylon sphaerocarpum DC.
  635. Memecylon sphaerothyrsum O. Schwartz
  637. Memecylon steenisii Bakh. f.
  639. Memecylon stenophyllum Merr.
  641. Memecylon stolzii Gilg ex Engl.
  643. Memecylon strumosum Naudin
  645. Memecylon strychnoides Gilg; nom. illeg.
  647. Memecylon subcaudatum Merr.
  649. Memecylon subcordifolium Bakh. f.
  651. Memecylon subcuneatum H. Perrier
  653. Memecylon subfurfuraceum Merr.
  655. Memecylon subramanii A.N. Henry
  657. Memecylon subsessile H. Perrier
  659. Memecylon subtrinervium Miq.
  661. Memecylon sumatrense Bakh. f.
  663. Memecylon superbum A. Fern. & R. Fern.
  665. Memecylon sylvaticum Thwaites
  667. Memecylon symplociforme Merr.
  669. Memecylon talbotianum Brandis
  671. Memecylon tamifolium Gilg ex Engl.
  673. Memecylon tayabense Merr.
  675. Memecylon teitense Wickens
  677. Memecylon tenuipes Merr.
  679. Memecylon terminale Dalzell
  681. Memecylon terminaliflorum Elmer
  683. Memecylon tetrapterum Cogn.
  685. Memecylon thomsonii Thwaites
  687. Memecylon thouarsianum Naudin
  689. Memecylon thouvenotii Danguy
  691. Memecylon thwaitesii Cogn.
  693. Memecylon tinctorium sensu Blanco
  695. Memecylon tirunelvelicum Murugan, Manickam & Sundaresan
  697. Memecylon toamasinense Jacq.-Fél.
  699. Memecylon torrei A. Fern. & R. Fern.
  701. Memecylon torricellense Lauterb.
  703. Memecylon tricolor Craib
  705. Memecylon trinerve DC.
  707. Memecylon tsaratananense Jacq.-Fél.
  709. Memecylon uapacoides Jacq.-Fél.
  711. Memecylon ulopterum DC.
  713. Memecylon umbellatum sensu Merr.
  715. Memecylon urceolatum Cogn.
  717. Memecylon utericarpum Jacq.-Fél.
  719. Memecylon vaccinioides Jacq.-Fél.
  721. Memecylon varians Thwaites
  723. Memecylon verruculosum Brenan
  725. Memecylon virescens Hook. f.
  727. Memecylon viride Hutch. & Dalziel; nom. inval.
  729. Memecylon vitiense A. Gray
  731. Memecylon walkeri Hook. ex C.B. Clarke; nom. nud.
  733. Memecylon wallichii Ridl.
  735. Memecylon wightii Thwaites
  737. Memecylon xiphophyllum R.D. Stone
  739. Memecylon zambeziense A. Fern. & R. Fern.
  741. Memecylon zenkeri Gilg