Pacific chupare The Pacific chupare or Pacific whiptail stingray , Styracura pacifica , is a species of stingray in the family Potamotrygonidae . It has been reported off the Pacific coast of Central America from Oaxaca, Mexico to Costa Rica , and also around the Galapagos Islands . It is usually found in shallow water on soft silty or muddy flats; it is unknown whether this species is tolerant of low salinity like the related chupare stingray . Leonard Compagno doubted the taxonomic validity of this species in his 1999 Checklist of Living Elasmobranchs .Appearance This stingray attains a maximum known length of and a disk width of. It has a rounded pectoral fin disk and a broadly angled snout with a small protuberance at the tip. The tail lacks fin folds but has a low ventral keel . The dorsal surface of the body and tail are covered with rough dermal denticles . There are large tubercles with four radial ridges on the shoulder region. A venomous spine is present on the tail.Parasites and behavior Known parasites of this species include the nematode Echinocephalus janzeni and the cestodes Acanthobothroides pacificus and Rhinebothrium geminum . Reproduction is ovoviviparous .Taxonomy The Pacific chupare and the chupare stingray in the Atlantic are believed to be sister species , together referred to as the "amphi-American Himantura ". The two species are morphologically similar and share four-ridged shoulder tubercles. Based on the details of the mandibular musculature and articulation, the amphi-American Himantura are hypothesized to be most closely related to the river stingrays in the family Potamotrygonidae, rather than to Indo-Pacific Himantura species. This has given rise to the theory that both the amphi-American Himantura and the river stingrays are descended from euryhaline ancestors living along the northern coast of South America prior to the formation of the Isthmus of Panama . This interpretation was initially disputed , as parasitological evidence suggests that the river stingrays are most closely related to Pacific Urobatis stingrays. In 2016, a major review of "Himantura " based on morphology and molecular evidence confirmed the position of the Pacific chupare, and it was moved to the genus Styracura in the family Potamotrygonidae.
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