Polyphaga


Polyphaga is the largest and most diverse suborder of beetles. It comprises 144 families in 16 superfamilies, and displays an enormous variety of specialization and adaptation, with over 350,000 described species, or approximately 90% of the beetle species so far discovered.
Key characteristics of Polyphaga are that the hind coxa, does not divide the first and second abdominal/ventral plates which are known as sternites. Also, the notopleural suture is not present.

Etymology

The name of polyphaga is derived from two Greek words: poly-, meaning 'many', and phagein, meaning 'to eat', so the suborder is called the “eaters of many things”.

Classification

The five infraorders are:
The internal classification of Polyphaga involves several superfamilies or series, whose constituents are relatively stable, although some smaller families are allocated to different clades by different authors. Large superfamilies include Hydrophiloidea, Staphylinoidea, Scarabaeoidea, Buprestoidea, Byrrhoidea, Elateroidea, and Bostrichoidea.
The infraorder Cucujiformia includes the vast majority of phytophagous beetles, united by cryptonephric Malpighian tubules of the normal type, a cone ommatidium with open rhabdom, and lack of functional spiracles on the eighth abdominal segment. Constituent superfamilies of Cucujiformia are Cleroidea, Cucujoidea, Tenebrionoidea, Chrysomeloidea, and Curculionoidea. Evidently adoption of a phytophagous lifestyle correlates with taxon diversity in beetles, with Cucujiformia, especially weevils, forming a major radiation.