Pterostichinae
Pterostichinae is a subfamily of ground beetles. It belongs to the advanced harpaline assemblage, and if these are circumscribed sensu lato as a single subfamily, Pterostichinae are downranked to a tribe Pterostichini. However, as the former Pterostichitae supertribe of the Harpalinae as loosely circumscribed does seem to constitute a lineage rather distinct from Harpalus, its core group is here considered to be the present subfamily and the Harpalinae are defined more narrowly.
They are usually mid-sized and rather stout ground beetles. Coloration is typically dark and without conspicuous patterns, but often with a strong sheen like polished metal. They are widely distributed and inhabit a wide range of terrestrial habitats. Unlike the more basal ground beetles which only eat small animals, the Pterostichinae include a large proportion of omnivorous or even herbivorous taxa.
Systematics
This group includes the following tribes and genera:Tribe Abacetini Chaudoir, 1872
- Abacetus Dejean, 1828
- Abacidus LeConte, 1873
- Aristopus Laferte-Senectere, 1853
- Celioinkosa Straneo, 1951
- Chlaeminus Motschulsky, 1865
- Colpodichius Straneo, 1952
- Cosmodiscus Sloane, 1907
- Cyrtomoscelis Chaudoir, 1874
- Distrigidius Jeannel, 1948
- Ecnomolaus Bates, 1892
- Haptoderidius Straneo, 1975
- Haptoderodes Straneo, 1986
- Holconotus Schmidt-Goebel, 1846
- Inkosa Peringuey, 1926
- Mateuellus Deuve, 1990
- Metabacetus Bates, 1892
- Metaxellus Straneo, 1960
- Metaxys Chaudoir, 1857
- Novillidius Straneo, 1941
- Oodinkosa Straneo, 1939
- Ophonichius Straneo, 1942
- Pediomorphus Chaudoir, 1878
- Pioprosopus Tschitscherine, 1899
- Pollicobius Vinson, 1939
- Prostalomus Basilewsky, 1950
- Pseudabacetus Burgeon, 1935
- Pterostillichus Straneo, 1949
- Rhagadillius Straneo, 1951
- Tiferonia Darlington, 1962
- Trachelocyphoides Alluaud, 1934
- Trachelocyphus Tschitscherine, 1900
- Catapiesis Solier, 1835
- Homalomorpha Brulle, 1835
- Chaetodactyla Tschitscherine, 1897
- Cnemalobus Guerin-Meneville, 1838
- Brachidius Chaudoir, 1852
- Cratocerus Dejean, 1829
- Oxyglychus Straneo, 1938
- Abacaelostus Straneo, 1952
- Andrewesinulus Straneo, 1938
- Apsidocnemus Alluaud, 1936
- Barylaus Liebherr, 1985
- Caecocaelus Straneo, 1949
- Caelostomus W.S. MacLeay, 1825
- Camptogenys Tschitscherine, 1899
- Capabatus Csiki, 1930
- Crenulostrigus Straneo, 1942
- Cyrtolaus Bates, 1882
- Dactyleurys Tschitscherine, 1899
- Dactylinius Straneo, 1941
- Diachipteryx Alluaud, 1926
- Diceromerus Chaudoir, 1872
- Dromistomus Jeannel, 1948
- Drymonaxus Straneo, 1941
- Eurystomis Chaudoir, 1878
- Feostoma Straneo, 1941
- Hemitelestus Alluaud, 1895
- Hoplizomenus Chaudoir, 1872
- Leleuporites Straneo, 1960
- Madapelmus Dajoz, 1985
- Monodryxus Straneo, 1942
- Pachycaecus Straneo, 1971
- Pachyroxochus Straneo, 1942
- Platyxythrius Straneo, 1942
- Stegazopteryx Will, 2004
- Stomonaxellus Tschitscherine, 1901
- Strigomerodes Straneo, 1939
- Strigomerus Chaudoir, 1872
- Trichillinus Straneo, 1938
- Microcheila Brulle, 1834
- Buderes Murray, 1857
- Hyperectenus Alluaud, 1935
- Hyperion Castelnau, 1834
- Megamorio Chaudoir, 1880
- Morion Latreille, 1810
- Morionidius Chaudoir, 1880
- Moriosomus Motschulsky, 1864
- Platynodes Westwood, 1846
- Stereostoma Murray, 1857
- Abacillius Straneo, 1949
- Abacillodes Straneo, 1988
- Abacoleptus Fauvel, 1903
- Abacomorphus Chaudoir, 1878
- Abacophrastus Will, 2011
- Abacops Tschitscherine, 1902
- Abaris Dejean, 1831
- Abax Bonelli, 1810
- Abropus Waterhouse, 1842
- Acanthoferonia Moore, 1965
- Adrimus Bates, 1871
- Allotriopus Bates, 1882
- Antarctiola Straneo, 1977
- Apsaustodon Tschitscherine, 1901
- Argutoridius Chaudoir, 1876
- Aristochroa Tschitscherine, 1898
- Aristochroodes Marcillac, 1993
- Aulacopodus Britton, 1940
- Basilewskya Straneo, 1948
- Blennidus Motschulsky, 1865
- Blenniventer Straneo, 1986
- Bothynoproctus Tschitscherine, 1900
- Camptoscelis Dejean, 1828
- Castelnaudia Tschitscherine, 1890
- Catadromus W.S. Macleay, 1825
- Cedrorum Borges & Serrano, 1993
- Cephalostichus Straneo, 1977
- Cerabilia Castelnau, 1867
- Chaetauchenium Tschitscherine
- Chalcochrous Chaudoir, 1838
- Conchitella Moore, 1962
- Cophosomorpha Tschitscherine, 1890
- Cratoferonia Tschitscherine, 1902
- Cratogaster Blanchard, 1843
- Cyclotrachelus Chaudoir, 1838
- Cynthidia Chaudoir, 1873
- Darodilia Castelnau, 1867
- Eucamptognathus Chaudoir, 1837
- Euchroa Brulle, 1834
- Eudromus Klug, 1835
- Eumara Tschitscherine, 1901
- Euryabax Fauvel, 1903
- Euryaptus Bates, 1892
- Feroniola Tschitscherine, 1900
- Feronista Moore, 1965
- Gastrellarius Casey, 1918
- Gastrogmus Sloane, 1915
- Gourlayia Britton, 1964
- Haplobothynus Tschitscherine, 1901
- Haploferonia Darlington, 1962
- Harpostomus Chaudoir, 1856
- Henrotius Jeannel, 1953
- Holcaspis Chaudoir, 1865
- Homalonesiota Maindron, 1908
- Hybothecus Chaudoir, 1874
- Kuschelinus Straneo, 1963
- Leiradira Castelnau, 1867
- Lesticus Dejean, 1828
- Liopasa Tschitscherine, 1901
- Litarthrum Sloane, 1915
- Lobobrachus Sharp, 1885
- Lophoglossus LeConte, 1852
- Loxandrus LeConte, 1852
- Loxodactylus Chaudoir, 1865
- Loxogenius Sloane, 1907
- Marsyas Putzeys, 1845
- Mecynognathus W.J.MacLeay, 1873
- Megadromus Motschulsky, 1865
- Meropalpus Tschitscherine, 1900
- Metius Curtis, 1839
- Metoncidus Bates, 1871
- Microcephalus Dejean, 1828
- Mireius Straneo, 1984
- Molopidius Jeannel, 1942
- Molopinus Jeannel, 1948
- Molops Bonelli, 1810
- Myas Sturm, 1826
- Nebrioferonia Straneo, 1939
- Nelidus Chaudoir, 1878
- Neoferonia Britton, 1940
- Neotalus Will, 2002
- Nirmala Andrewes, 1930
- Notabax Moore, 1976
- Notolestus Sloane, 1894
- Notonomus Chaudoir, 1862
- Nurus Motschulsky, 1866
- Oberthueria Vuillet, 1911
- Ogmophora Tschitscherine, 1896
- Oribazus Chaudoir, 1874
- Orthomus Chaudoir, 1838
- Oscadytes Lagar Mascaro, 1975
- Oxycrepis Reiche, 1843
- Pachymorphus Chaudoir, 1838
- Pachythecus Chaudoir, 1874
- Paniestichus Will, 2011
- Paranurus Tschitscherine, 1901
- Pareuryaptus Dubault, Lassalle & Roux, 2008
- Parhypates Motschulsky, 1866
- Pedius Motschulsky, 1850
- Percolaus Bates, 1882
- Percus Bonelli, 1810
- Peyrieraselus Deuve, 1981
- Phaenaulax Tschitscherine, 1898
- Piesmus LeConte, 1852
- Platycaelus Blanchard, 1843
- Platysmodes Fauvel, 1903
- Plocamostethus Britton, 1940
- Poecilinus Jeannel, 1948
- Poecilus Bonelli, 1810
- Prosopogmus Chaudoir, 1865
- Psegmatopterus Chaudoir, 1878
- Pseudabarys Chaudoir, 1873
- Pseudoceneus Tschitscherine, 1891
- Pterostichus Bonelli, 1810
- Rhabdotus Chaudoir, 1865
- Rhytisternus Chaudoir, 1865
- Sarticus Motschulsky, 1865
- Secatophus Castelnau, 1867
- Setalidius Chaudoir, 1878
- Setalimorphus Sloane, 1895
- Setalis Castelnau, 1867
- Simodontus Chaudoir, 1843
- Speluncarius Reitter, 1886
- Speomolops Patrizi, 1955
- Sphodrosomus Perroud & Montrouzier, 1864
- Stereocerus Kirby, 1837
- Steropanus Fairmaire, 1888
- Sthenocranion Tschitscherine, 1899
- Stolonis Motschulsky, 1866
- Stomis Clairville, 1806
- Straneostichus Sciaky, 1994
- Styracoderus Chaudoir, 1874
- Taphoxomimus Straneo, 1975
- Tapinopterus Schaum, 1858
- Teratotarsa Tschitscherine, 1893
- Teropha Castelnau, 1867
- Trichosternus Chaudoir, 1865
- Trigonognatha Motschulsky, 1858
- Trigonotoma Dejean, 1828
- Trirammatus Chaudoir, 1838
- Tropidocerus Chaudoir, 1878
- Wahlbergiana Bousquet, 2002
- Xenion Tschitscherine, 1902
- Zariquieya Jeannel, 1924
- Zeodera Castelnau, 1867
- Zeopoecilus Sharp, 1886
- Amara Bonelli, 1810
- Pseudamara Lindroth, 1968
- Zabrus Clairville, 1806