Pyrgini The Pyrgini are a tribe in the skipper butterfly subfamily Pyrginae . Formerly, when only four tribes of Pyrginae were recognized, the Pyrgini contained the largest number of genera among these. But this overly wide delimitation has since turned out to be paraphyletic . One of the traditional Pyrginae tribes, the Eudamini , had to be raised to subfamily rank as Eudaminae . Some genera now in the Eudaminae were placed in the Pyrgini in earlier times. In addition, a number of additional tribes - Achlyodidini , Carcharodini and Erynnini - are now usually recognized again. These are close relatives of the Pyrgini sensu stricto , and may just as well be included in them as they used to be: together they do still form a monophyletic group . Most authors prefer to keep them separate however, as each is an apomorphic and biogeographically distinct lineage of the Pyrgini sensu lato . Of these newly recognized tribes , the Achlyodidini are closest to the Pyrgini, but conspicuously differ from the latter in their unusually-shaped wings .Genera The genera of the Pyrgini sensu stricto are: Anisochoria Antigonus Carrhenes Celotes - streaky skippers Clito Cornuphallus Diaeus Eracon Heliopetes Heliopyrgus Onenses Paches Plumbago Pseudodrephalys Pyrgus - grizzled skippers , "checkered skippers" Spioniades Systasea Timochreon Trina Xenophanes Zobera Zopyrion Godman & Salvin , The genus Polyctor is also considered part of the tribe.Footnotes
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