Rubus
Rubus is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, with 250–700 species.
Raspberries, blackberries, and dewberries are common, widely distributed members of the genus. Most of these plants have woody stems with prickles like roses; spines, bristles, and gland-tipped hairs are also common in the genus. The Rubus fruit, sometimes called a bramble fruit, is an aggregate of drupelets. The term "cane fruit", or "cane berry", applies to any Rubus species or hybrid which is commonly grown with supports such as wires or canes, including raspberries, blackberries, and hybrids such as loganberry, boysenberry, marionberry and tayberry. The stems of such plants are also referred to as canes.
Overview
Most species are hermaphrodites, Rubus chamaemorus being an exception.The blackberries, as well as various other Rubus species with mounding or rambling growth habits, are often called brambles. However, this name is not used for those like the raspberry that grow as upright canes, or for trailing or prostrate species, such as most dewberries, or various low-growing boreal, arctic, or alpine species.
The generic name means blackberry in Latin and was derived from the word ruber, meaning "red".
The scientific study of brambles is known as "".
Examples of the hundreds of species of Rubus include:
- Rubus aboriginum – garden dewberry
- Rubus abundus
- Rubus aculifer
- Rubus adenocaulis
- Rubus adenophorus
- Rubus adenotrichos
- Rubus adjacens
- Rubus alaskensis
- Rubus allegheniensis – Allegheny blackberry
- Rubus alnifolius
- Rubus alpinus
- Rubus alter
- Rubus alumnus
- Rubus amplior
- Rubus andrewsianus
- Rubus apogaeus
- Rubus aptatus
- Rubus arcticus – Arctic raspberry
- Rubus arcuans
- Rubus arenicola
- Rubus argutus
- Rubus arizonensis – Arizona dewberry
- Rubus armeniacus – Himalayan blackberry
- Rubus arvensis
- Rubus audax
- Rubus azuayensis
- Rubus baileyanus
- Rubus bartonianus
- Rubus bellobatus
- Rubus bicknellii
- Rubus biflorus
- Rubus biformispinus
- Rubus bifrons
- Rubus bigelovianus
- Rubus blanchardianus
- Rubus boyntonii
- Rubus burnhamii
- Rubus bushii
- Rubus caesius – European dewberry
- Rubus canadensis – Canadian blackberry
- Rubus canescens
- Rubus centralis
- Rubus chamaemorus – Cloudberry
- Rubus chloocladus
- Rubus clarus
- Rubus cockburnianus
- Rubus conanicutensis
- Rubus concameratus
- Rubus corchorifolius
- Rubus coriifolius
- Rubus coronarius
- Rubus coreanus – Bokbunja
- Rubus costaricanus
- Rubus crataegifolius
- Rubus cubitans
- Rubus cuneifolius – sand blackberry
- Rubus curtipes
- Rubus cymosus
- Rubus dalibarda
- Rubus deamii
- Rubus defectionis
- Rubus deliciosus
- Rubus densissimus
- Rubus depavitus – Aberdeen dewberry
- Rubus discolor
- Rubus dissimilis
- Rubus domingensis
- Rubus durescens
- Rubus durus
- Rubus eggersii
- Rubus elegantulus
- Rubus ellipticus
- Rubus emeritus
- Rubus eriocarpus
- Rubus exeter
- Rubus exsularis
- Rubus fagifolius
- Rubus fecundus
- Rubus felix
- Rubus ferrugineus
- Rubus flagellaris – northern dewberry
- Rubus flavinanus
- Rubus floricomus
- Rubus florulentus
- Rubus fraternalis
- Rubus frondisentis
- Rubus frondosus
- Rubus fruticosus agg. – blackberry
- Rubus fryei
- Rubus furtivus
- Rubus geniculatus
- Rubus geoides – Patagonian raspberry
- Rubus glabratus
- Rubus glandicaulis
- Rubus glaucifolius – San Diego raspberry
- Rubus glaucus
- Rubus gnarus
- Rubus grimesii
- Rubus griseus
- Rubus gulosus
- Rubus hancinianus
- Rubus hanesii
- Rubus harmonicus
- Rubus hawaiensis
- Rubus hayata-koidzumii
- Rubus heterophyllus
- Rubus hirsutus
- Rubus hispidoides
- Rubus hispidus – swamp dewberry
- Rubus hochstetterorum – Azorean blackberry
- Rubus humistratus
- Rubus huttonii
- Rubus hypolasius
- Rubus ictus
- Rubus idaeus – European red raspberry
- Rubus illecebrosus
- Rubus immanis
- Rubus impar
- Rubus inclinis
- Rubus inferior
- Rubus iniens
- Rubus insons
- Rubus insulanus
- Rubus invisus – upland dewberry
- Rubus irasuensis
- Rubus ithacanus
- Rubus jacens
- Rubus jamaicensis
- Rubus junceus
- Rubus kelloggii
- Rubus kennedyanus
- Rubus laciniatus – cutleaf evergreen blackberry
- Rubus laegaardii
- Rubus lambertianus
- Rubus largus
- Rubus lasiococcus – roughfruit berry
- Rubus latens
- Rubus lawrencei
- Rubus leucodermis – whitebark raspberry or western raspberry
- Rubus leviculus
- Rubus liebmannii
- Rubus linkianus
- Rubus longii
- Rubus macraei
- Rubus macrogongylus
- Rubus macrophyllus
- Rubus macvaughii
- Rubus maniseesensis
- Rubus meracus
- Rubus michiganensis
- Rubus mirus
- Rubus miser
- Rubus missouricus
- Rubus mollior
- Rubus moluccanus
- Rubus montensis
- Rubus multifer
- Rubus multiformis
- Rubus multispinus
- Rubus navus
- Rubus nefrens
- Rubus nelsonii
- Rubus nebulosus
- Rubus neomexicanus
- Rubus nepalensis
- Rubus nigerrimus
- Rubus nivalis
- Rubus niveus
- Rubus notatus
- Rubus novanglicus
- Rubus noveboracus
- Rubus novocaesarius
- Rubus obsessus
- Rubus obvius
- Rubus occidentalis – black raspberry
- Rubus odoratus – flowering raspberry
- Rubus oklahomus
- Rubus originalis
- Rubus ortivus
- Rubus ostryifolius
- Rubus paganus
- Rubus palmeri
- Rubus paludivagus
- Rubus parcifrondifer
- Rubus parlinii
- Rubus particeps
- Rubus particularis
- Rubus parviflorus – thimbleberry
- Rubus parvifolius – small-leaf bramble
- Rubus pascuus
- Rubus pedatus
- Rubus pensilvanicus – Pennsylvania blackberry
- Rubus pergratus
- Rubus permixtus
- Rubus pernagaeus
- Rubus persistens
- Rubus perspicuus
- Rubus pervarius
- Rubus philadelphicus
- Rubus philyrophyllus
- Rubus phoenicolasius – wineberry
- Rubus pittieri
- Rubus plexus
- Rubus plicatifolius
- Rubus plicatus
- Rubus plus
- Rubus porteri
- Rubus positivus
- Rubus prestonensis
- Rubus pringlei
- Rubus probabilis
- Rubus probativus
- Rubus probus
- Rubus prosper
- Rubus provincialis
- Rubus pubescens – dwarf red blackberry
- Rubus pugnax
- Rubus pumilus
- Rubus quaesitus
- Rubus racemiger
- Rubus randolphiorum
- Rubus recurvans
- Rubus recurvicaulis
- Rubus reflexus – rusty-hair raspberry
- Rubus regionalis
- Rubus repens
- Rubus riograndis
- Rubus roribaccus
- Rubus rosa
- Rubus rosarius
- Rubus roseus
- Rubus rosifolius
- Rubus rossbergianus
- Rubus russeus
- Rubus rydbergianus
- Rubus saltuensis
- Rubus sapidus
- Rubus saxatilis – stone bramble
- Rubus scambens
- Rubus scandens
- Rubus sceleratus
- Rubus schiedeanus
- Rubus schoolcraftianus
- Rubus segnis
- Rubus semisetosus
- Rubus setosus
- Rubus severus
- Rubus sewardianus
- Rubus sieboldii
- Rubus sierrae
- Rubus signatus
- Rubus sons
- Rubus spectabilis – salmonberry
- Rubus spectatus
- Rubus steelei
- Rubus stipulatus
- Rubus strigosus – American red raspberry
- Rubus superbus
- Rubus suppar
- Rubus suus
- Rubus tardatus
- Rubus thibetanus
- Rubus tholiformis
- Rubus tomentosus
- Rubus trichomallus
- Rubus tricolor
- Rubus trifidus – Japanese blackberry
- Rubus trifrons
- Rubus trilobus
- Rubus trivialis
- Rubus tuerckheimii
- Rubus tygartensis
- Rubus ucetanus
- Rubus uhdeanus
- Rubus ulmifolius
- Rubus uniformis
- Rubus ursinus – trailing blackberry
- Rubus uvidus
- Rubus variispinus
- Rubus velox
- Rubus verae-crucis
- Rubus vermontanus
- Rubus vestitus
- Rubus vigil
- Rubus vigilis
- Rubus vigoratus
- Rubus vulcanicola
- Rubus vulgaris
- Rubus weatherbyi
- Rubus whartoniae
- Rubus wheeleri
- Rubus wisconsinensis
Hybrid berries
The term "hybrid berry" is often used collectively for those fruits in the genus Rubus which have been developed mainly in the USA and UK in the last 130 years. As Rubus species readily interbreed and are apomicts, the parentage of these plants is often highly complex, but is generally agreed to include cultivars of blackberries, and raspberries.The hybrid berries include:-
- Loganberry R. × loganobaccus, a spontaneous cross between R. ursinus 'Aughinbaugh' and R. idaeus 'Red Antwerp'
- Boysenberry a cross between R. idaeus and R. × loganobaccus
- Olallieberry a cross between loganberry and youngberry, themselves both hybrid berries.
- Veitchberry a cross between R. fruticosus and R. idaeus
- Skellyberry, a cross between R. invisus and R. phoenicolasius
- Marionberry now thought to be a blackberry cultivar R. 'Marion'
- Silvanberry, R. 'Silvan', a cross between R. 'Marion' and boysenberry
- Tayberry, another blackberry/raspberry cross
- Tummelberry, R. 'Tummel', from the same Scottish breeding programme as tayberry
- Hildaberry, a tayberry/boysenberry cross discovered by an amateur grower
- Youngberry, a complex hybrid of raspberries, blackberries, and dewberries
Scientific classification
Rubus species have a basic chromosome number of seven. Polyploidy from the diploid to the tetradecaploid is exhibited.
Some treatments have recognized dozens of species each for what other, comparably qualified botanists have considered single, more variable species. On the other hand, species in the other Rubus subgenera are generally distinct, or else involved in more routine one-or-a-few taxonomic debates, such as whether the European and American red raspberries are better treated as one species or two.
Molecular data have backed up classifications based on geography and chromosome number, but following morphological data, such as the structure of the leaves and stems, do not appear to produce a phylogenetic classification.
The classification presented below recognizes 13 subgenera within Rubus, with the largest subgenus in turn divided into 12 sections. Representative examples are presented, but many more species are not mentioned here. A comprehensive 2019 study found subgenera Orobatus and Anoplobatus to be monophyletic, while all other subgenera to be paraphyletic or polyphyletic.
Fossil record
The genus has a likely North American origin, with the oldest fossils known from the Eocene, and expansion into Eurasia and elsewhere during the Miocene. Fossil seeds from the early Miocene of Rubus have been found in the Czech part of the Zittau Basin.Many fossil fruits of †Rubus laticostatus, †Rubus microspermus and †Rubus semirotundatus have been extracted from bore hole samples of the Middle Miocene fresh water deposits in Nowy Sacz Basin, West Carpathians, Poland.