Tyranni


The Tyranni are a clade of passerine birds that includes more than 1,000 species, the large majority of which are South American. It is named after the type genus Tyrannus.
These have a different anatomy of the syrinx musculature than the oscines, hence its common name of suboscines. The available morphological, DNA sequence, and biogeographical data, as well as the fossil record, agree that these two major passerine suborders are evolutionarily distinct clades.

Systematics

The suborder Tyranni is divided into two infraorders: the Eurylaimides and the Tyrannides. The New Zealand wrens in the family Acanthisittidae are placed in a separate suborder Acanthisitti.
The Eurylaimides contain the Old World suboscines – mainly distributed in tropical regions around the Indian Ocean – and a single American species, the sapayoa:
Phylogenetic relationships of the Eurylaimides based on Oliveros et al. :

The Tyrannides contain all the suboscines from the Americas, except the broad-billed sapayoa. The families listed here are those recognised by the International Ornithologists' Union.
Phylogenetic relationships of the Tyrannides based on Oliveros et al. :
This group has been separated into three parvorders by Sibley & Ahlquist. However, DNA-DNA hybridization has shown to be not very well suited to reliably resolve the suboscine phylogeny. It was eventually determined that there was a simple dichotomy between the antbirds and allies, and the tyrant-flycatchers and allies. Given that the "parvorder" arrangement originally advanced is certainly obsolete - more so if the Eurylaimides are elevated to a distinct suborder - it would be advisable to rank the clades as superfamilies, or if the broadbill group is considered a separate suborder, as infraorders. In the former case, the name Furnarioidea would be available for the tracheophones, whereas "Tyrannoidea", the "bronchophone" equivalent, has not yet been formally defined.
In the latter case, the tracheophones would be classified as "Furnariides",
while the Tyrannides would be restricted to the tyrant-flycatchers and other "bronchophone" families.
The tracheophones contain the Furnariidae, Thamnophilidae, Formicariidae, and Conopophagidae. The tyrant-flycatcher clade includes the namesake family, the Tityridae, the Cotingidae, and the Pipridae.