Lowndes was born in Winslow, Buckinghamshire, the son of Robert Lowndes and his second wife, Elizabeth FitzWilliam. His father was descended from the Lowndes of Overton in Cheshire, but his grandfather had moved south to Buckinghamshire. Other relatives had settled in South Carolina. Lowndes was educated at the free school in Buckingham. He joined HM Treasury as a clerk. He acquired, Chesham's Bury Manor in 1687
Political career
Lowndes was returned unopposed as a Member of Parliament for Seaford in Sussex, a "limb" of the Cinque Ports, at the 1695 general election. He was returned unopposed for Seaford for every election until 1715. He served as chairman of the Committee of Ways and Means, becoming known throughout Britain as "Ways and Means Lowndes". Lowndes also became Secretary to the Treasury in 1695. His Report containing an essay for the amendment of the silver coins, written during the crisis of 1695, was answered by John Locke, whose views on the reform of the currency prevailed. He became wealthy as a result of holding office in the Treasury. In 1700, he built Winslow Hall in Winslow, Buckinghamshire. Around this period Lowndes was painted twice by Sir Godfrey Kneller, and by contemporary painterRichard Philips whose portrait is in the collection of the Bank of England. Lowndes originated the funded system and rose to great power and influence in Parliament. In recognition of his service, Queen Anne conferred upon him the office of Auditor of the Land Revenue for life, in reversion to his sons, with an augmentation to his coat of arms. In 1712 Lowndes rebuilt Chesham's manor house, The Bury, immediately to the south of St Mary's Church, Chesham. The building still stands today, and currently serves as an office building. At the 1715 general election Lowndes was returned unopposed as MP for St Mawes in Cornwall in the first Parliament of King George I, but stood unsuccessfully for Westminster in 1722. Shortly afterwards, he was returned in a by-election on 27 October 1722 as MP for East Looe, also in Cornwall, after that constituency was vacated by Horace Walpole when he decided to stand for Great Yarmouth instead. In 1723 Lowndes bought the freehold reversion of leasehold property he owned in St. James's and Knightsbridge, in areas now known as Lowndes Square and Lowndes Street. The expression "Take care of the pence, and the pounds will take care of themselves" is attributed to him.
Death and legacy
Lowndes' death was announced in the House of Commons by Walpole, saying The House had lost a very useful Member, and the public as able and honest a servant as ever the Crown had.
Lowndes married four times:
Elizabeth Harsnett, daughter of Sir Roger Harsnett
Lowndes had children with each of his wives, 25 children in all. His offspring were also fruitful: one son had 16 children, including four sets of twins in four years; a grandson had 10 children. He was succeeded by his son Richard, who inherited Winslow Hall and became both High Sheriff and MP for Buckinghamshire. William Lowndes, a son by his third wife, followed him as an Auditor of His Majesty's Court of Exchequer in the 1760s.