Yaguarasaurus Yaguarasaurus is an extinct genus of mosasauroid from the Late Cretaceous period of Colombia , South America . The remains discovered were defined as a new genus and species of mosasaurid, Yaguarasaurus columbianus , by the Colombian paleontologist María Páramo , former director of the Museo de Geología José Royo y Gómez of INGEOMINAS in Bogotá . The first fossils remains of this animal suggested a cranial length of and a total length of ; an additional skull that measures long implies a larger size. This reptile is a member of the family of marine lizards Mosasauridae characteristic of Middle and Upper Cretaceous , with global distribution, but in South America known only through isolated remains. This mosasaur discovered in Yaguará , was at the moment of discovery the most complete material known in South America.Etymology The remains were found in a limestone bed of the La Frontera Formation , member of the Villeta Group , near Yaguará, Huila , in a site called Cueva Rica. Its name means "Yaguará lizard of Colombia".Phylogeny In the initial descriptions of Yaguarasaurus , it was classified as a member of the subfamily Plioplatecarpinae ; the analysis of Polcyn and Bell showed that it was a distant relative of the advanced mosasaurids and were a close relative of Russellosaurus coheni and Tethysaurus nopscai , a group that could be a basal clade to the division between Tylosaurinae and Plioplatecarpinae subfamilies, called Russellosaurina . After the discovery of the basal mosasauroid of Hungary Pannoniasaurus by Makádi et al . 2012 it became clear that Yaguarasaurus and the rest of the "russellosaurines" form a clade of basal amphibian mosasaurs related with the aigialosaurs, a grouping called the subfamily Tethysaurinae ; the cladogram below follow the analysis of Makádi et al. de 2012: Palci et al. describe a new Russellosaurine, Romeosaurus , and created subfamily Yaguarasaurinae, sister group of Tethysaurinae
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