Human Y-chromosome DNA haplogroup
In human genetics, a human Y-chromosome DNA haplogroup is a haplogroup defined by mutations in the non-recombining portions of DNA from the Y chromosome. Many people within a haplogroup share a type of mutation called single-nucleotide polymorphisms.
The human Y-chromosome accumulates roughly two mutations per generation. Y-DNA haplogroups represent major branches of the Y-chromosome phylogenetic tree that share hundreds or even thousands of mutations unique to each haplogroup.
The Y-chromosomal most recent common ancestor is the most recent common ancestor from whom all currently living humans are descended patrilineally. Y-chromosomal Adam is estimated to have lived roughly 236,000 years ago in Africa. By examining other bottlenecks most Eurasian men are descended from a man who lived 69,000 years ago. Other major bottlenecks occurred about 50,000 and 5,000 years ago and subsequently most Eurasian/non-African men can trace their ancestry back to four ancestors who lived 50,000 years ago.
Naming convention
Y-DNA haplogroups are defined by the presence of a series of Y-DNA SNP markers. Subclades are defined by a terminal SNP, the SNP furthest down in the Y-chromosome phylogenetic tree. The Y Chromosome Consortium developed a system of naming major Y-DNA haplogroups with the capital letters A through T, with further subclades named using numbers and lower case letters. YCC shorthand nomenclature names Y-DNA haplogroups and their subclades with the first letter of the major Y-DNA haplogroup followed by a dash and the name of the defining terminal SNP.Y-DNA haplogroup nomenclature is changing over time to accommodate the increasing number of SNPs being discovered and tested, and the resulting expansion of the Y-chromosome phylogenetic tree. This change in nomenclature has resulted in inconsistent nomenclature being used in different sources. This inconsistency, and increasingly cumbersome longhand nomenclature, has prompted a move towards using the simpler shorthand nomenclature. In September 2012, Family Tree DNA provided the following explanation of its changing Y-DNA haplogroup nomenclature to individual customers on their Y-DNA results pages :
Phylogenetic structure
;Phylogenetic tree of Y-DNA haplogroupsMajor Y-DNA haplogroups
Haplogroups A and B
Haplogroup A is the NRY macrohaplogroup from which all modern paternal haplogroups descend. It is sparsely distributed in Africa, being concentrated among Khoisan populations in the southwest and Nilotic populations toward the northeast in the Nile Valley. BT is a subclade of haplogroup A, more precisely of the A1b clade, as follows:- Haplogroup A
- * Haplogroup A00
- * Haplogroup A0
- * Haplogroup A1
- ** Haplogroup A1a
- ** Haplogroup A1b
- *** Haplogroup A1b1a1
- *** Haplogroup A1b1b
- *** Haplogroup BT
- **** Haplogroup B
- **** Haplogroup CT
Haplogroup CT (P143)
Haplogroup C (M130)
- Haplogroup C Found in Asia, Oceania, and North America
- * Haplogroup C1
- ** Haplogroup C1a
- *** Haplogroup C1a1 Found with low frequency in Japan
- *** Haplogroup C1a2 Found with low frequency in Europe, Armenians, Algeria, and Nepal
- ** Haplogroup C1b
- *** Haplogroup C1b1
- **** Haplogroup C1b1a
- ***** Haplogroup C1b1a1 Found with low frequency in South Asia, Southwest Asia, and northern China
- ***** Haplogroup C1b1a2
- ****** Haplogroup C1b1a2a Found among Lebbo' people in Borneo, Indonesia
- ****** Haplogroup C1b1a2b Found among Han Chinese, Dai people, Murut people, Malay people, and Aeta people
- ***** Haplogroup C1b1a3 Found with low frequency in Saudi Arabia and Iraq
- **** Haplogroup C1b1b Found among Dusun people
- *** Haplogroup C1b2
- *** Haplogroup C1b3
- **** Haplogroup C1b3a Found in Indonesia, New Guinea, Melanesia, Micronesia, and Polynesia
- **** Haplogroup C1b3b Found among the indigenous peoples in Australia
- * Haplogroup C2 Found throughout Eurasia and North America, but especially among Mongols, Kazakhs, Tungusic peoples, Paleosiberians, and Na-Dené-speaking peoples
Haplogroup D (CTS3946)
- Haplogroup D
- * Haplogroup D1 Found in Japan, China, the Andaman Islands
- ** Haplogroup D1a
- *** Haplogroup D1a1
- **** Haplogroup D1a1a Found mainly in Tibetans, Qiangic peoples, Yi, and Hmong-Mien peoples
- **** Haplogroup D1a1b Found mainly in Tibetans, Qiangic peoples, Naxi, and Turkic peoples
- *** Haplogroup D1a2 Found mainly in Japan
- *** Haplogroup D1a3 Found in Andamanese peoples
- ** Haplogroup D1b Found in Mactan Island, Philippines
- * Haplogroup D2 Found in Nigeria, Saudi Arabia and Syria
Haplogroup E (M96)
- Haplogroup E Found in Africa and parts of the Middle East and Europe
- * Haplogroup E1
- ** Haplogroup E1a formerly E1
- ** Haplogroup E1b
- *** Haplogroup E1b1 ; formerly E3
- **** Haplogroup E1b1a
- ***** Haplogroup E1b1a1 Found in Africa, especially among Niger–Congo-speaking populations.; formerly E3a
- ***** Haplogroup E1b1a2 Found in Africa, especially in Ethiopia among Omotic-speaking populations.; formerly E3*
- **** Haplogroup E1b1b
- ***** Haplogroup E1b1b1 Found in Horn of Africa, North Africa, the Middle East, and Europe ; formerly E3b
- * Haplogroup E2
Haplogroup F (M89)
F xG,H,I,J,K is rare in modern populations and peaks in South Asia, especially Sri Lanka. It also appears to have long been present in South East Asia; it has been reported at rates of 4–5% in Sulawesi and Lembata. One study, which did not comprehensively screen for other subclades of F-M89, found that Indonesian men with the SNP P14/PF2704, comprise 1.8% of men in West Timor, 1.5% of Flores 5.4% of Lembata 2.3% of Sulawesi and 0.2% in Sumatra. F* has been reported among 10% of males in Sri Lanka and South India, 5% in Pakistan, as well as lower levels among the Tamang people, and in Iran. F1, F2 and F3 are all highly rare and virtually exclusive to regions/ethnic minorities in Sri Lanka, India, Nepal, South China, Thailand, Burma, and Vietnam. In such cases, however, the possibility of misidentification is considered to be relatively high and some may belong to misidentified subclades of Haplogroup GHIJK.
Haplogroup G (M201)
originated some 48,000 years ago and its most recent common ancestor likely lived 26,000 years ago in the Middle East. It spread to Europe with the Neolithic Revolution.It is found in many ethnic groups in Eurasia; most common in the Caucasus, Iran, Anatolia and the Levant. Found in almost all European countries, but most common in Gagauzia, southeastern Romania, Greece, Italy, Spain, Portugal, Tyrol, and Bohemia with highest concentrations on some Mediterranean islands; uncommon in Northern Europe.
G-M201 is also found in small numbers in northwestern China and India, Bangladesh, Pakistan, Sri Lanka, Malaysia, and North Africa.
- Haplogroup G1
- Haplogroup G2
- * Haplogroup G2a
- ** Haplogroup G2a1
- ** Haplogroup G2a2
- ** Haplogroup G2a3
- *** Haplogroup G2a3a
- *** Haplogroup G2a3b
- **** Haplogroup G2a3b1
- ** Haplogroup G2b
- ** Haplogroup G2c
- *** Haplogroup G2c1
Haplogroup H (M69)
However, H2 is present in Europe since the Neolithic and H1a1 spread westward in the Medieval era with the migration of the Roma people.
Haplogroup I (M170)
Haplogroup I is found mainly in Europe and the Caucasus.- Haplogroup I1 Nordid/Nordic Euopids Found mainly in northern Europe
- Haplogroup I2 Dinarid/Dinaric Europids Found mainly in Balkans, southeast Europe and Sardinia save for I2B1 which is found at a moderate frequency in Western, Central, and Northern Europe.
Haplogroup J (M304)
- Haplogroup J* is rare outside the island of Socotra.
- * Haplogroup J1 Semitid/Bedouinid Arabids is associated with Northeast Caucasian peoples in Dagestan and Semitic languages speaking people in the Middle East, Ethiopia, and North Africa and also found in Mediterranean Europe in smaller frequencies much like haplogroup T.
- * Haplogroup J2 Syrid/Nahrainid Arabids is found mainly in the Semitic-speaking peoples, Anatolia, Greece, the Balkans, Italy, Iran, the Caucasus, South Asia, and Central Asia.
Haplogroup K (M9)
K – that is, K*, K2c, K2d or K2e – is found mainly in Melanesia, Aboriginal Australians, India, Polynesia and Island South East Asia.
Haplogroups L and T (K1)
is found in South Asia, Central Asia, South-West Asia, and the Mediterranean.Haplogroup T is found at high levels in the Horn of Africa, parts of South Asia, the Middle East, and the Mediterranean. T-M184 is also found in significant minorities of Sciaccensi, Stilfser, Egyptians, Omanis, Sephardi Jews, Ibizans, and Toubou. It is also found at low frequencies in other parts of the Mediterranean and South Asia.
Haplogroup K2 (K-M526)
The only living males reported to carry the basal paragroup K2* are indigenous Australians. Major studies published in 2014 and 2015 suggest that up to 27% of Aboriginal Australian males carry K2*, while others carry a subclade of K2.Haplogroups K2a, K2a1, NO & NO1
Haplogroup N
is found through northern Eurasia, especially among speakers of the Uralic languages.Haplogroup N possibly originated in eastern Asia and spread both northward and westward into Siberia, being the most common group found in some Uralic-speaking peoples.
Haplogroup O
Haplogroup O is found with its highest frequency in East Asia and Southeast Asia, with lower frequencies in the South Pacific, Central Asia, South Asia, and islands in the Indian Ocean.- Haplogroup O1
- * Haplogroup O1a Found in eastern,central and southern Mainland China, Taiwan, and Southeast Asia, especially among Austronesian and Tai–Kadai peoples
- * Haplogroup O1b
- ** Haplogroup O1b1 Found in Japan, southern China, Taiwan, Southeast Asia, and the Indian subcontinent, especially among Austroasiatic- and Tai–Kadai-speaking peoples, Malays, and Indonesians
- ** Haplogroup O1b2 Found in Japan, Korea, Manchuria, and Southeast Asia
- Haplogroup O2 Found throughout East Asia, Southeast Asia, and Austronesia including Polynesia
Haplogroups K2b1, M & S
Its primary subclades are two major haplogroups:
- Haplogroup S also known as K2b1a: found in the highlands of Papua New Guinea and;
- Haplogroup M also known as K2b1b: found in New Guinea and Melanesia.
Haplogroup P (K2b2)
P*, P1* and P2 are found together only on the island of Luzon, in The Philippines. In particular, P* and P1* are found at significant rates among members of the Aeta people of Luzon. While, P1* is now more common among living individuals in Eastern Siberia and Central Asia, it is also found at low levels in mainland South East Asia and South Asia. Considered together, these distributions tend to suggest that P* emerged from K2b in South East Asia.
P1 is also the parent node of two primary clades:
- Haplogroup Q and;
- Haplogroup R. These share the common marker M45 in addition to at least 18 other SNPs.
Haplogroup R : found in Europe, West Asia, Central Asia, and South Asia
Haplogroup Q M242
Q is defined by the SNP M242. It is believed to have arisen in Central Asia approximately 32,000 years ago. The subclades of Haplogroup Q with their defining mutation, according to the 2008 ISOGG tree are provided below. ss4 bp, rs41352448, is not represented in the ISOGG 2008 tree because it is a value for an STR. This low frequency value has been found as a novel Q lineage in Indian populationsThe 2008 ISOGG tree
- Q
- * Q*
- * Q1
- ** Q1*
- ** Q1a
- *** Q1a*
- *** Q1a1 Found with low frequency among Bhutanese, Dungans, Han Chinese, Japanese, Koreans, Mongolians, Naxi, and Tibetans
- *** Q1a2 Found at low to moderate frequency among some populations of Southwest Asia, Central Asia, and Siberia
- *** Q1a3
- **** Q1a3* Found at low frequency in Pakistan, India, and Tibet
- **** Q1a3a Typical of indigenous peoples of the Americas
- ***** Q1a3a*
- ***** Q1a3a1 Found among some indigenous peoples of South America, such as the Ticuna and the Wayuu
- ***** Q1a3a2
- ***** Q1a3a3
- *** Q1a4
- *** Q1a5
- *** Q1a6 Found in a significant minority of Yemeni Jews
- ** Q1b Found at low frequency among samples of Hazara and Sindhis
Haplogroup R (M207)
R1a is associated with the proto-Indo-Iranian and Balto-Slavic peoples, and is now found primarily in Central Asia, South Asia, and Eastern Europe.
Haplogroup R1b is the dominant haplogroup of Western Europe and also found sparsely distributed among various peoples of Asia and Africa. Its subclade R1b1a2 is the haplogroup that is most commonly found among modern Western European populations, and has been associated with the Italo-Celtic and Germanic peoples.
- Haplogroup R1 Found throughout western Eurasia
- * Haplogroup R1a Found in Central Asia, South Asia, and Central, Northern and Eastern Europe, Balkans
- * Haplogroup R1b Found in Western Europe, West Asia, Central Asia, North Africa, and northern Cameroon
- Haplogroup R2 Found in South Asia, Caucasus, Central Asia, and Eastern Europe
Chronological development of haplogroups