List of canids
is a family of mammals in the order Carnivora, which includes domestic dogs, wolves, coyotes, foxes, jackals, dingoes, and many other extant and extinct dog-like mammals. A member of this family is called a canid; all extant species are a part of a single subfamily, Caninae, and are called canines. They are found on all continents except Antarctica, having arrived independently or accompanied human beings over extended periods of time. Canids vary in size, including tails, from the 2 meter gray wolf to the 46 cm fennec fox. Population sizes range from the Falkland Islands wolf, extinct since 1876, to the wolf, whose domestic dog subspecies has a worldwide population of over 1 billion. The body forms of canids are similar, typically having long muzzles, upright ears, teeth adapted for cracking bones and slicing flesh, long legs, and bushy tails. Most species are social animals, living together in family units or small groups and behaving cooperatively. Typically, only the dominant pair in a group breeds, and a litter of young is reared annually in an underground den. Canids communicate by scent signals and vocalizations. One canid, the domestic dog, entered into a partnership with humans at least 14,000 years ago and today remains one of the most widely kept domestic animals.
The 13 genera and 37 species of Caninae are primarily split into two tribes: Canini, which includes 9 genera and 20 species, comprising the wolf-like Canina subtribe and the South American Cerdocyonina subtribe; and Vulpini, the fox-like canids, comprising 3 genera and 14 species. Not included in either tribe is the Urocyon genus, which includes 2 species, mainly comprising the gray fox and believed to be basal to the family. In addition to the extant Caninae, Canidae comprises two extinct subfamilies designated as Hesperocyoninae and Borophaginae. Extinct species have also been placed into Caninae, in both extant and extinct genera; at least 80 extinct Caninae species have been found, as well as over 70 species in Borophaginae and nearly 30 in Hesperocyoninae, though due to ongoing research and discoveries the exact number and categorization is not fixed. The earliest canids found belong to Hesperocyoninae, and are believed to have diverged from the existing Caniformia suborder around 37 million years ago.
Conventions
Conservation status codes listed follow the International Union for Conservation of Nature Red List of Threatened Species. Range maps are provided wherever possible; if a range map is not available, a description of the canid's range is provided. Ranges are based on the IUCN red list for that species unless otherwise noted. All extinct species or subspecies listed alongside extant species went extinct after 1500 CE, and are indicated by a dagger symbol "".Classification
The family Canidae consists of 35 extant species belonging to 12 genera and divided into 194 extant subspecies, as well the extinct genus Dusicyon, comprising two extinct species, and 13 extinct wolf subspecies, which are the only canid species to go extinct since prehistoric times. This does not include hybrid species or extinct prehistoric species. Modern molecular studies indicate that the 13 genera can be grouped into 3 tribes or clades.Subfamily Caninae
- Tribe Canini
- *Subtribe Canina
- **Genus Canis: seven species
- **Genus Cuon: one species
- **Genus Lycaon: one species
- *Subtribe Cerdocyonina
- **Genus Atelocynus: one species
- **Genus Cerdocyon: one species
- **Genus Chrysocyon: one species
- **Genus Dusicyon: two species
- **Genus Lycalopex: six species
- **Genus Speothos: one species
- Tribe Vulpini
- *Genus Nyctereutes: one species
- *Genus Otocyon: one species
- *Genus Vulpes: twelve species
- Genus Urocyon: two species
Canids
Subfamily Caninae
Tribe Canini
Tribe Vulpini
Urocyon
Prehistoric canids
In addition to extant canids, a number of prehistoric species have been discovered and classified as a part of Canidae. Morphogenic and molecular phylogenic research has placed them within the extant subfamily Caninae as well as the extinct subfamilies Hesperocyoninae and Borophaginae. Within Caninae, prehistoric species have been placed into both extant genera and separate extinct genera.The generally accepted classification of extinct canid species is primarily based for Hesperocyoninae on work by Xiaoming Wang, curator of terrestrial mammals at the Natural History Museum of Los Angeles County, and on work by Wang and zoologists Richard H. Tedford and Beryl E. Taylor for Borophaginae and Caninae. The species and classifications listed below are all from these works; exceptions due to more recently-described species are also listed with citations. Not all of these classifications are universally accepted, and alternate classifications for species are noted below. Where available, the approximate time period the species was extant is given in millions of years before the present, based on data from the Paleobiology Database. All listed species are extinct; where a genus, subtribe, or tribe within Caninae comprises only extinct species, it is indicated with a dagger symbol.
Subfamily Caninae
- Tribe Canini
- * Subtribe Canina
- ** Genus Canis
- *** C. antonii
- *** C. apolloniensis
- *** C. armbrusteri
- *** C. arnensis
- *** C. brevicephalus
- *** C. cedazoensis
- *** C. chihliensis
- *** C. cipio
- *** C. dirus
- *** C. edwardii
- *** C. etruscus
- *** C. ferox
- *** C. gezi
- *** C. leilhardi
- *** C. lepophagus
- *** C. longdanensis
- *** C. mosbachensis
- *** C. nehringi
- *** C. othmani
- *** C. palmidens
- *** C. variabilis
- ** Genus Cynotherium
- *** C. sardous
- ** Genus Eucyon
- *** E. adoxus
- *** E. davisi
- *** E. intrepidus
- *** E. monticinensis
- *** E. odessanus
- *** E. skinneri
- *** E. zhoui
- ** Genus Lycaon
- *** L. magnus
- *** L. sekowei
- ** Genus Mececyon
- *** M. trinilensis
- ** Genus Megacyon
- *** M. merriami
- ** Genus Nurocyon
- *** N. chonokhariensis
- ** Genus Xenocyon
- *** X. africanus
- *** X. antonii
- *** X. falconeri
- *** X. lycaonoides
- * Subtribe Cerdocyonina
- ** Genus Cerdocyon
- *** C. avius
- *** C. ensenadensis
- *** C. texanus
- ** Genus Nyctereutes
- *** N. abdeslami
- *** N. donnezani
- *** N. megamastoides
- *** N. sinensis
- *** N. terblanchei
- *** N. tingi
- ** Genus Protocyon
- *** P. orcesi
- *** P. scagliarum
- *** P. troglodytes
- ** Genus Speothos
- *** S. pacivorus
- ** Genus Theriodictis
- *** T. floridanus
- Tribe Vulpini
- * Genus Metalopex
- ** M. bakeri
- ** M. macconnelli
- ** M. merriami
- * Genus Prototocyon
- ** P. curvipalatus
- ** P. recki
- * Genus Vulpes
- ** V. alopecoides
- ** V. angustidens
- ** V. beihaiensis
- ** V. chikushanensis
- ** V. galaticus
- ** V. praecorsac
- ** V. praeglacialis
- ** V. riffautae
- ** V. skinneri
- ** V. stenognathus
- ** V. qiuzhudingi
- Urocyon
- * Genus Urocyon
- ** U. minicephalus
- ** U. progressus
- Basal Caninae
- * Genus Leptocyon
- ** L. delicatus
- ** L. douglassi
- ** L. gregorii
- ** L. leidyi
- ** L. matthewi
- ** L. mollis
- ** L. tejonensis
- ** L. vafer
- ** L. vulpinus
- Unclassified
- * Genus Protemnocyon
- ** P. inflatus
Subfamily Borophaginae
- Tribe Borophagini
- * Genus Cormocyon
- ** C. copei
- ** C. haydeni
- * Genus Desmocyon
- ** D. matthewi
- ** D. thomsoni
- * Genus Euoplocyon
- ** E. brachygnathus
- ** E. spissidens
- * Genus Metatomarctus
- ** M. canavus
- * Genus Microtomarctus
- ** M. conferta
- * Genus Protomarctus
- ** P. optatus
- * Genus Psalidocyon
- ** P. marianae
- * Genus Tephrocyon
- ** T. rurestris
- * Subtribe Aelurodontina
- ** Genus Aelurodon
- *** A. asthenostylus
- *** A. ferox
- *** A. mcgrewi
- *** A. montanensis
- *** A. stirtoni
- *** A. taxoides
- ** Genus Tomarctus
- *** T. brevirostris
- *** T. hippophaga
- * Subtribe Borophagina
- ** Genus Borophagus
- *** B. diversidens
- *** B. dudleyi
- *** B. hilli
- *** B. littoralis
- *** B. orc
- *** B. parvus
- *** B. pugnator
- *** B. secundus
- ** Genus Carpocyon
- *** C. compressus
- *** C. limosus
- *** C. robustus
- *** C. webbi
- ** Genus Epicyon
- *** E. aelurodontoides
- *** E. haydeni
- *** E. saevus
- ** Genus Paratomarctus
- *** P. euthos
- *** P. temerarius
- ** Genus Protepicyon
- *** P. raki
- * Subtribe Cynarctina
- ** Genus Cynarctus
- *** C. crucidens
- *** C. galushai
- *** C. marylandica
- *** C. saxatilis
- *** C. voorhiesi
- *** C. wangi
- ** Genus Paracynarctus
- *** P. kelloggi
- *** P. sinclairi
- Tribe Phlaocyonini
- * Genus Cynarctoides
- ** C. acridens
- ** C. emryi
- ** C. gawnae
- ** C. harlowi
- ** C. lemur
- ** C. luskensis
- ** C. roii
- * Genus Phlaocyon
- ** P. achoros
- ** P. annectens
- ** P. latidens
- ** P. leucosteus
- ** P. mariae
- ** P. marslandensis
- ** P. minor
- ** P. multicuspus
- ** P. taylori
- ** P. yatkolai
- Basal Borophaginae
- * Genus Archaeocyon
- ** A. falkenbachi
- ** A. leptodus
- ** A. pavidus
- * Genus Otarocyon
- ** O. cooki
- ** O. macdonaldi
- * Genus Oxetocyon
- ** O. cuspidatus
- * Genus Rhizocyon
- ** R. oregonensis
Subfamily Hesperocyoninae
- Genus Cynodesmus
- * C. martini
- * C. thooides
- Genus Caedocyon
- * C. tedfordi
- Genus Ectopocynus
- * E. antiquus
- * E. intermedius
- * E. siplicidens
- Genus Enhydrocyon
- * E. basilatus
- * E. crassidens
- * E. pahinsintewkpa
- * E. stenocephalus
- * E. sectorius
- Genus Hesperocyon
- * H. coloradensis
- * H. gregarius
- Genus Mesocyon
- * M. brachyops
- * M. coryphaeus
- * M. temnodon
- Genus Osbornodon
- * O. brachypus
- * O. fricki
- * O. iamonensis
- * O. renjiei
- * O. scitulus
- * O. sesnoni
- * O. wangi
- Genus Paraenhydrocyon
- * P. josephi
- * P. robustus
- * P. wallovianus
- Genus Philotrox
- * P. condoni
- Genus Prohesperocyon
- * P. wilsoni
- Genus Sunkahetanka
- * S. geringensis